Kolora lynneae, Raven, 2015

Kolora lynneae View in CoL sp. nov.

( Figs 48b–d View FIGURE 48 , 52a–f View FIGURE 52 . Map 27)

Material. Queensland: holotype ♂, Gayundah Ck , Hinchinbrook I, 18°22'S, 146°13'E, rainforest, 7–15 Nov 1984, G. Monteith, G. Thompson, D. Cook, QM S28409 View Materials . Paratypes GoogleMaps : Queensland: 1♂, Mission Beach (S2), 17°52'S, 146°05'E, rainforest, pitfall, 4 Dec 1995 – 2 Jan 1996, M. Cermak, QM S39287 View Materials ; 1♂, Innisfail , 17°31'S, 146°01'E, 30 May 1997, J. Hasenpusch, AM KS97253 ; allotype 1♀, Tinaroo Ck, Mareeba, AM KS52061 GoogleMaps .

Diagnosis. Differs from all other species in dark brown sash centrally down carapace; males have only the apical embolus irregularly rugose, not spiralled.

Etymology. The species epithet is in honour of the daughter, Lynne Rawlinson, of collection manager and artist, Wendy Hebron.

Description holotype male QM S28409 View Materials

Carapace 3.28 long, 2.00 wide. Abdomen 3.20 long, 1.44 wide. Total length 6.5.

Colour: carapace brown yellow with broad sash from and including chelicerae, full width of caput, narrowing slightly in front of fovea with irregular margins back to posterior margin; palp, leg I yellowish; II yellowish except for brown flanks on femora; III: femora brown, pallid dorsally and ventrally; IV missing; abdomen brown with pile of short golden feathery hairs and distal cap of pallid hair; sternum, coxae and trochanters yellowish. Carapace: uniform pile of short white feathery hairs; fovea shallow, open; long golden brown hairs directed forward along cuticle in front, beside and slightly behind eyes. Chelicerae: both margins with two large teeth set midlength, retromargin also with one smaller distal; fang shield small, distinct, rounded. Maxillae: slight ectal concavity medially. Sternum with two additional bristles medially. Legs: both legs IV missing, tufts larger on III than on I; all leg femora with feathery hairs. Spines: leg I: fe: p3d5r2; pa 0; ti v2.2; me v2.1.1 all long. II: fe: p3d4r3; pa 0; ti v2.2; me v2.1.1 all long. III: fe: p3d3r4; pa 0; ti p2r2v2; me p2r2v2.1.1. Palp: fe p1d1; pa and tibia all long bristles. Palp: as for K. cooloola sp. nov. but paired claw-spine present on cymbium tip. Abdomen: long narrow dorsal scute for two-thirds length, genital scute only weakly sclerotised; mid-dorsal with saddle; ventral scute long narrow fused to tracheal scute; colulus and spiracle distinct, unsclerotised.

Allotype female AM KS52061 . As male but:

Carapace 3.48 long, 2.06 wide. Abdomen 4.46 long, 2.34 wide. Total length, 9.1. Colour: carapace orange brown with broad brown medial stripe for length; abdomen abdomen matt black with transverse bar at midlength and large pallid area terminally; dorsal scute small, confined to anterior slope; palps yellow brown; pedal femora basally brown, I distally yellow brown, II–IV, brown. Cheliceral fang mound low with line of ca. 12 long curved serrated bristles for cheliceral width. Spines: tibiae and metatarsi I, II with 2 pairs long thin spines ventrally. Tracheal scute small, lenticular. Epigyne: CO angular, oval, just over one diameter apart.

Distribution and Habitat. Known from rainforest at Hinchinbrook Island, and Mission Beach, Innisfail and Mareeba, northeastern Queensland.

Leichhardteus Raven & Baehr, 2013

Leichhardteus Raven & Baehr View in CoL , in Baehr & Raven, 2013: 340. Type species by original designation, Leichhardteus conopalpis Baehr & Raven, 2013 View in CoL .

Diagnosis. Females differ from those of Battalus in lacking the strong setae on the fang shield and the posterior lobe of the spermathecae is not smooth walled but folded (e.g., Fig. 62b View FIGURE 62 ); males differ from those of Battalus and Corinnomma in the inornate embolus tip, the weak or absent paracymbial spine and in all, save for L. strzelecki sp. nov. and L.kroombit Baehr & Raven, 2013 ( Fig. 59e View FIGURE 59 ), in the presence of a mound or spur retrobasally on the palpal femur (e.g., Figs 53c View FIGURE 53 , 61d View FIGURE 61 ).

Description. Females lack strong setae on fang shield, a low plate with weak setae. Lateral profile of carapace ( Fig. 2b View FIGURE 2 ): caput arched anteriorly, sloping through slight arch in shallow slope from fovea along posterior declivity; highest point just behind eye group; eyes set behind clypeus edge. Retrolateral face of large chelicerae smooth. Eight eyes in two rows of similar width; front row recurved, back row procurved ( Fig. 16d View FIGURE 16 ). AME much larger than PLE larger than ALE larger than PME. Maxillae basally truncate, ectal margin slightly convex, with rounded apex and pallid truncation on anterior inner corners. Labium as wide as long, shield-shaped. Sternum shield-shaped without precoxal sclerites and not continuous with pedicel. Legs: I about same size as IV; leg lengths 4123, coxae entally rounded with slight retrobasal process. RCH small, distinct. Trochanters symmetrically notched about twice as wide as deep, similar on all legs. No feathery hairs on legs; covered mostly with bristles, not hairs. Tarsi cylindrical for length. Tibiae I and II with 3–4 pairs of spines ventrally; metatarsi I and II with two pairs ventrally. Female palp: tarsus with one distal pair of spines ventrally; no spines ventrally on femur. Trichobothria on tarsi and metatarsi few, short, distal; tibiae with cluster of trichae basally. Claws: two, enclosed in short dense tufts; claws and tufts on legs I and II very small, lower than diameter of tarsi; claws longer with few small teeth on legs III and IV. Scopula a fringe of short hairs on ventral tarsi I–IV and distally on all metatarsi. Abdomen: males with small to large oval dorsal scute (e.g., Fig. 57d View FIGURE 57 ); venter only with genital scute; females with small oval scute dorsally and tracheal scute small, ovoid with rounded ridge. Epigyne ( Figs 56c View FIGURE 56 , 58c View FIGURE 58 ): externally a domed plate, longer than wide with lateral subdistal crescentic hoods opening posteriorly, from which copulatory duct sweeps behind large dumbbell lobes of spermathecae. Paragenital brackets present or absent.

Males. Palp: femur with spur ventrally just distal of midpoint, varying from absent, subtle ( Fig. 59e View FIGURE 59 ) to hornlike (e.g., Figs 53c View FIGURE 53 , 61d View FIGURE 61 ); tibia cylindrical but with distinct glabrous saddle distoventrally for half of prolateral edge ( Fig. 59f View FIGURE 59 ) where small process interlocks with ovoid depression on opposed cymbial edge ( Fig. 55b–e View FIGURE 55 ); cymbium with sinuous retrolateral margin ( Fig. 55c, d View FIGURE 55 ) and conical apical tip; paracymbial spine absent or vestigial ( Figs 57b View FIGURE 57 , 58b View FIGURE 58 ) to long, slender ( Fig. 56b View FIGURE 56 ); typical simple castianeirine bulb with enlarged basal part of tegulum; sperm duct folds back strongly toward embolus then reflexes back sharply; sclerotised collar, round base of short embolus with reflexed flattened apex; no cymbial scopula.

Distribution and Habitat. Coastal areas of eastern, southern and Western Australia. In eastern Australia, the spiders are found in rainforest, semievergreen vine thicket, and open forest. In southern and western Australia, the spiders are found in mallee, heath and open forest.

Species Included: Leichhardteus albofasciatus Baehr & Raven, 2013 , L. badius Baehr & Raven, 2013 , L. bimaculatus Baehr & Raven, 2013 , L. conopalpis Baehr & Raven, 2013 , L. evschlingeri sp. nov., L. garretti Baehr & Raven, 2013 , L. kroombit Baehr & Raven, 2013 , L. reinhardi Baehr & Raven, 2013 , L. strzelecki sp. nov., L. yagan sp. nov. and L. terriirwinae Baehr & Raven, 2013 .

Remarks. Baehr & Raven (2013) diagnosed the genus by the femoral prong on the palp of males (e.g., Figs 53c View FIGURE 53 , 61d View FIGURE 61 ). However, save for the prong, which becomes marginal in some species, L. strzelecki sp. nov. fits well into Leichhardteus . Species descriptions are expanded to allow comparison to other genera.

Key to species of Leichhardteus View in CoL

Males

1. Palpal femur ventrally with high conical or pointed prong ( Fig. 57c View FIGURE 57 )............................................. 2 Palpal femur ventrally with low mound ( Fig. 55f View FIGURE 55 ) or prong absent ( Fig. 59e View FIGURE 59 )....................................... 6

2(1). Femoral prong with abrupt basal angle junction with femur ( Fig. 57c View FIGURE 57 )............................................ 3 Femoral prong with gradual curving junction with femur ( Baehr & Raven, 2013, fig. 7I)...................... L. garretti View in CoL

3(2). Paracymbial spine small but distinct ( Figs 55e View FIGURE 55 , 56b View FIGURE 56 ).......................................................... 4 Paracymbial spine absent or a weak spine (e.g., Fig. 57b View FIGURE 57 )...................................................... 5

4(3). Femoral prong clearly triangular ( Baehr & Raven, 2013, fig. 10G).................................... L. terriirwinae Femoral View in CoL prong slender, almost digitiform ( Fig. 57c View FIGURE 57 )............................................... L. evschlingeri View in CoL

5(3). Femoral prong low as wide, basally as high, retrolateral ( Fig. 61d View FIGURE 61 )......................................... L. yagan Femoral View in CoL prong high ( Fig. 53c View FIGURE 53 )................................................................. L. conopalpis View in CoL

6(1). Paracymbial spine absent ( Fig. 54b View FIGURE 54 )....................................................................... 7 Paracymbial spine small but distinct ( Fig. 60f View FIGURE 60 )............................................................... 8

7(6). Femoral prong absent or very indistinct........................................................... L. strzelecki Femoral View in CoL prong distinct ( Baehr & Raven, 2013, fig. 5H)................................................. L. badius View in CoL

8(6). Four pairs (distal may be much shorter) of paired spines on ventral tibiae I and II; femoral prong low but well demarcated ( Baehr & Raven, 2013, Fig. 9I View FIGURE 9 )........................................................................... 9 Three pairs of paired spines on ventral tibiae I and II; femoral prong indistinct, subtle or not well demarcated ( Fig. 59e View FIGURE 59 )................................................................................................. L. kroombit View in CoL

9(8). Abdomen dorsally entirely black, without white spots ( Baehr & Raven, 2013, fig. 9A)...................... L. reinhardi View in CoL Abdomen with white spots or pallid median band........................................................... 10

10(9). Abdomen dark with two large white spots ( Baehr & Raven, 2013, fig. 6A)............................. L. bimaculatus View in CoL Abdomen dark with longitudinal irregular median band ( Fig. 55a View FIGURE 55 )................................... L. albofasciatus View in CoL

Females (females of L. bimaculatus View in CoL , L. evschlingeri View in CoL sp. nov., L. strzelecki View in CoL sp. nov. and L. terriirwinae View in CoL unknown) 1. Four pairs of spines on ventral tibiae I and II................................................................ 2

Three pairs of spines on ventral tibiae I and II............................................................... 6 2(1). Posterior spermathecal lobe smaller than anterior ( Fig. 53e View FIGURE 53 ).................................................... 3

Posterior spermathecal lobe similar size to anterior ( Fig. 62b View FIGURE 62 )................................................... 4 3(2). CO directed anterolaterally ( Baehr & Raven, 2013, fig. 7L)............................................. L. garretti View in CoL

CO directed posterolaterally ( Fig. 53d View FIGURE 53 )........................................................... L. conopalpis View in CoL 4(2). CO very small, 6 diameters apart ( Fig. 56c View FIGURE 56 )..................................................... L. albofasciatus View in CoL

CO larger, 2–5 diameters apart ( Fig. 62a View FIGURE 62 )................................................................... 5 5(4). CO 4–5 diameters apart ( Fig. 62a View FIGURE 62 ).................................................................. L. yagan View in CoL

CO 2–3 diameters apart ( Baehr & Raven, 2013, Fig. 9J View FIGURE 9 ).............................................. L. reinhardi View in CoL 6(1). CO relatively wider apart ( Fig. 54c View FIGURE 54 )................................................................ L. badius View in CoL

CO relatively closer ( Fig. 58c View FIGURE 58 )................................................................... L. kroombit View in CoL