Family
VERMETIDAE Rafinesque, 1815
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Tubispiracea – Deshayes, 1832-EncyMeth: table facing p. 553, as “les Tubispirés”, later latinized by Reeve (1882), but unavailable because not based on a generic name (ICZN Code Art. 11.7.1.1).
Tubispirata – Deshayes, 1861-DescrCoqFoss: 279-290. Emendation or spelling error for Tubispiracea Deshayes, 1832, q.v. Listed by Bieler & Petit (2011: 12).
anguillinus,
Serpulorbis
– Deshayes, 1861-DescrCoqFoss: 289-290, 9, pl. 9, figs 16-17, pl. 10, figs 5-6. — Type localities: Mercin-et-Vaux, Laon, Cuise-la-Motte, Mons-en-Laonnois & Hérouval. — Type age: Eocene (Ypresian). — Type species (OD) of
Anguillospira Cossmann, 1912
.
Vermicularia ( Anguillospira) anguillina (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105)
. Listed by Bieler & Petit (2011: 24). Anderson & Allmon (2023: 8) referred this taxon to the
Siliquariidae
.– Current status:
Anguillospira anguillina (Deshayes, 1861)
(Pacaud herein).
cancellatus
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,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 284-285, 9, pl. 9, fig. 8. — Type localities: Auvers-sur-Oise, Valmondois, Mary-sur-Marne, Jaignes, Crouy & Vendrest. — Type age: Eocene (Bartonian). — Current status:
Serpulorbis cancellatus Deshayes, 1861
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( Le Renard & Pacaud 1995: 105). Listed by Bieler & Petit (2011: 28).
clathratus
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,
Serpulorbis
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–Deshayes, 1861-DescrCoqFoss: 286-287, 9, pl. 9, figs 9-10. — Type localities: Auvers-sur-Oise, Vendrest, Crouy, Acy-en-Multien, Betz, Caumont & Mary-sur-Marne. — Type age: Eocene (Bartonian). Listed by Bieler & Petit (2011: 30). — Current status:
Serpulorbis clathratus Deshayes, 1861
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( Le Renard & Pacaud 1995: 105; Pacaud 2008: 57).
cristatus
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,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 287, 9, pl. 9, fig. 12, non
Vermetus cristatus Sandberger, 1860
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, nec Biondi, 1859. — Type localities: Auvers-sur-Oise & Mary-sur-Marne. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32398, syntype from Auvers-sur-Oise.
Vermetus excristatus Sacco, 1896
( Sacco 1896: 7), replacement name ( Bieler & Petit, 2011: 36).
Serpulorbis cristatus Deshayes, 1861
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( Le Renard & Pacaud 1995: 105; Pacaud 2008: 57). Listed by Bieler & Petit (2011: 33). — Current status: synonym of
Thylacodes excristatus ( Sacco, 1896)
.
laxatus
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,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 287-288, 9, pl. 9, fig. 15. — Type locality: Mary-sur-Marne. — Type age: Eocene (Bartonian). Listed by Bieler & Petit (2011: 44). — Current status:
Serpulorbis laxatus Deshayes, 1861
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( Le Renard & Pacaud 1995: 105; Pacaud 2008: 57).
masier
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,
Vermetus
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– Deshayes, 1843-HistNatAnim2: 65, ex Adanson ms; 1843-HistNatAnim3: 526. In the synonymy of
Serpula sipho Lamarck, 1818
. West Africa. — Type age: Recent. Listed by Bieler & Petit (2011: 44, 46). — Current status:
Thylacodes masier (Deshayes, 1843) (MolluscaBase)
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.
moerchi
,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 286, 9, as
“ morchi
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”, pl. 9, figs 21-22. — Type localities: Mercin-et-Vaux, Cuise-la-Motte & Laon. — Type age: Eocene (Ypresian). Stated to be named for Mörch, so automatically corrected (ICZN Code Art. 32.5).
Serpulorbis moerchi (Deshayes, 1861)
( Le Renard & Pacaud 1995: 105, as “
S. morchi
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”). Listed by Bieler & Petit (2011: 30). — Current status:
Thylacodes moerchi (Deshayes, 1861)
( Leroy et al. 2014: 25, pl. 21, figs 2-5).
ornatus
,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 285, 9, pl. 9, fig. 23, non
S. ornata (J. de C. Sowerby, 1850)
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[
Serpula
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]. — Type localities: Montmirail & Hautteville-Bocage. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32400, syntypes from Montmirail. Replaced as
Thylacodes deshayesi Newton, 1891
( Newton 1891: 217). However, the latter is a secondary homonym of
Serpulorbis deshayesi Mayer-Eymar, 1889
, which is also a
Thylacodes
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. Sacco (1896: 55) proposed as a replacement name
Vermetus deshayesianus
. A third replacement name,
Serpulorbis newtoni Glibert, 1962b
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( Glibert 1962b: 130), for
Serpulorbis ornatus Deshayes, 1861
, was unnecessary. All these taxa are
Thylacodes
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. The name
Serpula ornatus J. de C. Sowerby, 1850
, junior primary homonym of
Serpula ornata
I. Lea
, 1833, is now
Vermetus compsus Cossmann, 1892
( Cossmann 1892: 60-61), a replacement name. It is not a nomen nudum, as Bieler & Petit (2011: 30) asserted.
Serpulorbis ornatus Deshayes
, listed by Bieler & Petit (2011: 50). — Current status:
Thylacodes deshayesianus ( Sacco, 1896)
(Pacaud herein).
polygonus,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 290, 9, pl. 9, fig. 14. — Type localities: Chambors, Hérouval, Thiverval-Grignon & Valmondois. — Type age: Eocene (Lutetian-Bartonian).
Vermicularia ( Anguillospira) polygona (Deshayes, 1861)
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( Le Renard & Pacaud 1995: 105; Pacaud 2008: 49). Listed by Bieler & Petit (2011: 53). — Current status:
Anguillospira polygona (Deshayes, 1861)
(Pacaud herein).
porrectus,
Serpulorbis
– Deshayes, 1861-DescrCoqFoss: 288-289, 9, pl. 9, fig. 18. — Type localities: Acy-en-Multien, Crouy & La Ferté-sous-Jouarre. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32399, syntype from Acy-en-Multien.
Vermicularia ( Anguillospira) porrecta (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105)
. Listed by Bieler & Petit (2011: 53). — Current status:
Anguillospira porrecta (Deshayes, 1861)
(Pacaud herein).
semipedalis
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,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 288, 9, pl. 9, fig. 11. — Type localities: Valmondois, Auvers-sur-Oise, Crouy & Le Fayel. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32397, syntype from Auvers-sur-Oise. — Current status:
Serpulorbis semipedalis Deshayes, 1861
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( Le Renard & Pacaud 1995: 105). Listed by Bieler & Petit (2011: 58).
serpuloides
,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 289, 9, pl. 9, figs 19-20. — Type localities: Thiverval-Grignon, Mouchy-le-Châtel & Parnes. — Type age: Eocene (Lutetian).
Vermicularia ( Anguillospira) serpuloides (Deshayes, 1861)
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( Le Renard & Pacaud 1995: 105; Pacaud 2008: 49). Listed by Bieler & Petit (2011: 58). — Current status:
Thylacoides serpuloides (Deshayes, 1861)
(Pacaud herein).
strictus
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,
Serpulorbis
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– Deshayes, 1861-DescrCoqFoss: 285-286, 9, pl. 9, fig. 13. — Type locality: Auvers-sur-Oise. — Type age: Eocene (Bartonian). Listed by Bieler & Petit (2011: 61). — Current status:
Serpulorbis strictus (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105)
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.
turonensis
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,
Vermetus
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–Deshayes, 1850-TraitElem: 43, pl. 70, figs 14- 15. — Type locality: none given [ France]. — Type age: Miocene (Aquitanian).Listed by Bieler & Petit (2011: 66). — Current status:
Vermetus ( Lemintina) arenarius
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[Linnaeus, 1758;
Serpula
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]
turonensis Deshayes, 1850
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( Cossmann & Peyrot 1922: 81-82, pl. 3, figs 18- 20, as “1839”).
●
arenarius
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,
Vermetus
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– Deshayes, 1838-RegAnim: pl. 62, fig. 5; 1850-TraitElem: 43, pl. 70, fig. 16. — Type locality: none given. — Type age: probably Recent. Listed by Bieler & Petit (2011: 26) as a separate species. However, the plate explanation and caption with his original figure in his makes it clear that this name was labeled as “Desh” because it was a generic transfer of
Serpula arenaria Linnaeus, 1758
, to
Vermetus
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. Bieler & Petit (2011: 26, 70-71) regarded what they thought was Deshayes’s species as a nomen
dubium
. Deshayes’ figure in 1838 of an animal was copied from Philippi (1836: pl. 9, fig. 18a), which Philippi had had labeled
Vermetus gigas Bivona Bernardi, 1832
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. Deshayes evidently considered this taxon to be
a junior
synonym of Linnaeus’ species. His later figure in the Traité was only of a shell. This species is now listed as
Thylacodes arenarius (Linnaeus, 1758)
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; Linnaeus’ species is the senior synonym of the type species of
Thylacodes Guettard, 1770
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.
lispe, Vermet [ us] – See under Annelida
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.
tricarinatus
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,
Vermetus
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– Menke, 1844: 23, as “Deshayes, 1843”, nomen nudum.