Pterostichus ( Morphohaptoderus ) toledanoi Facchini & Sciaky, 2003

[11] Pterostichus ( Morphohaptoderus) toledanoi Facchini & Sciaky, 2003 View in CoL

Figs 4 E View Figure 4 , 5 G View Figure 5 , 23 View Figure 23

Pterostichus ( Morphohaptoderus) toledanoi Facchini & Sciaky, 2003: 11, ( holotype in CSF; type locality: Hubei: Shennongjia, 31 ° 24 ' 27 ' N, 110 ° 17 ' 20 ' E, 2500–2900 m). View in CoL

Type examined.

Holotype: Hubei • ♂, labeled “ CHINA, W-HUBEI, 2500–2900 m, Dashennongjia massif , 31 º 24 ’ – 27 ' N, 110 º 17 ’ – 20 ' E, lgt: 28.6. – 3.7. 95 L. & R. Businsky // HOLOTYPE, Pterostichus ( Morphohaptoderus) toledanoi n. sp., Det. Facchini & Sciaky 2000 ” [red label] ( CSF) . Paratypes: Hubei • 1 ♂, 1 ♀, the same data as holotype but labeled as paratype ( IZAS) .

Non-type material examined.

Hubei • 1 ♀, labeled “ Hubei Province, Shennongjia, Shennongding , N 31.4531 °, E 110.2681 °, under stone, 2890 m, 2022. VII. 27, Hongliang Shi & Ganyan Yang leg. ” [in Chinese] ( CBFU) GoogleMaps ; • 5 ♂, labeled “ Hubei, Shennongjia, Badongya , 2700 m, 1980. VII. 23, Peiyu Yu leg. ” [in Chinese] ; • 1 ♂, 3 ♀, labeled “ Jinhouling , Shennongjia, Hubei prov. 4. VIII. 2002, Li & Tang leg. ” ( IZAS) .

Diagnosis.

Medium-small-sized species for the subgenus ( BL = 7.5–7.8 mm); elytra with linear microsculpture in both sexes; antennomere 3 without accessory seta; pronotum nearly quadrate, lateral margins subparalleled behind middle; outer sulcus of pronotal basal fovea oblique to posterior margin of pronotum; elytral interval 3 with three setigerous pores, the basal one adjacent to stria 3, the other two adjacent to stria 2; fifth tarsomeres with 2–3 pairs of setae ventrally; male sternites simple; apical lamella of aedeagus large and capitate, apex truncated in lateral view.

Comparisons.

This species can be easily distinguished from nearly all other species of the subgenus Morphohaptoderus in China by its specific elytral chaetotaxy. Some species from Sichuan and one from Shaanxi ( P. ming Sciaky & Wrase, 1997 ) also have three setigerous pores on elytral interval 3. Nevertheless, in these species, the middle pore on the interval is always adjacent to striae 3, and the submentum has two setae on each side. In contrast, in P. toledanoi , the middle pore on the interval is adjacent to striae 2, while the submentum has only one seta on each side. This species can be confused only with the new species described below, P. xuanzhang sp. nov. Detailed comparisons between them will be presented under the description of the new species.

Description.

Very similar to P. xuanzhang sp. nov. externally. A complete redescription of external characters is unnecessary here, except for the following, which differ from P. xuanzhang sp. nov.

BL = 7.5–7.8 mm, BW = 3.1–3.2 mm; HW / PW = 0.58–0.59; tempora swollen behind eyes, about one-third length of eyes, eyL / tpL = 3.00–3.14. Pronotum subquadrate, PW / PL = 1.35–1.37, PBW / PW = 0.89–0.92, PAW / PBW = 0.67–0.71; posterior margin very strongly concaved at middle; outer sulcus of pronotal basal fovea distinctly oblique to the basal margin of pronotum, forming an angle at about 75 ° (Fig. 4 E View Figure 4 ). BW / EL = 0.69–0.72.

Median lobe of aedeagus (Fig. 23 A View Figure 23 ) relatively slender; in lateral view, ventral margin curved near basal third, nearly straight at middle, apex faintly bent ventrally; apical orifice relatively small, weakly turned to left side. Ventral surface with a straight sharp ridge extended from aedeagal base to nearly apex of apical lamella (Fig. 23 B View Figure 23 ). Apical lamella thick and capitate (Fig. 23 C View Figure 23 ), strongly bent upward, in lateral view, extreme apex truncated and slightly hooked at dorsal end; in dorsal view, apical lamella relatively long, in similar length as basal width ( AL / AW = 0.99–1.01), slightly bent to the left side. Right paramere short, apex rounded (Fig. 23 D View Figure 23 ).

Endophallus of male genitalia not studied. Gonocoxite 2 of female ovipositor relatively large, slightly crescent-shaped (Fig. 5 G View Figure 5 ), strongly obtuse apically; outer margin with two ensiform setae of normal size, length of the largest ensiform seta ca. 0.05 mm; dorsal margin with one ensiform seta near basal third, ridged between the ensiform seta and apex of gonocoxite 2, one nematiform seta located on the apical tenth of outer surface.

Distribution.

This species is only known from Shennongjia in Hubei and is distributed at high altitudes of 2500–2890 m.