Ocenebra schoenni (Hörnes, 1853)

Ocenebra schoenni (Hörnes, 1853)

Figs 13 View FIGURE 13 , 28F–H View FIGURE 28

* Murex Schönni Hörnes—Hörnes 1853: 235, pl. 24, fig. 12.

[ Murex craticulatus ] F. a pernutus De Greg.—De Gregorio 1885a: 245 [pro Murex craticulatus sensu Hörnes 1853 : pl. 24, fig. 12].

Murex ( Occenebra [sic]) Schönni M. Hoern.—Hoernes & Auinger 1885: 219.

Murex View in CoL ( Occenebra [sic]) Schönni Hoern.— Schaffer 1912: 143, pl. 50, figs 20–21.

Murex View in CoL ( Occenebra [sic]) crassilabiatus Hilb. View in CoL — Schaffer 1912: 143, pl. 50, fig. 13 [non Ocinebrinopsis crassilabiata ( Hilber 1879) ].

O [cinebrina]. schönni (Hörn.) — Sieber 1958: 146.

Ocinebrina schönni (Hörnes, 1856) — Báldi et al. 1964: 173, pl. 2, fig. 12.

[ Ocinebrina View in CoL ] schönni Hoernes, 1853 — Vokes 1971: 96.

Ocinebrina schönni (Hoernes, 1853) — Steininger et al. 1971: 399, pl. 10, figs 2–3.

Ocinebrina crassilabiata ( Hilber, 1879) View in CoL — Steininger et al. 1971: 399, pl. 10, fig. 4 [non Ocinebrinopsis crassilabiata ( Hilber 1879) ].

Ocinebrina schönni (Hörnes, 1856) — Báldi 1973: 289, pl. 37, fig. 3.

Type material. Lectotype (designated herein): NHMW 1852 View Materials /0012/0009a, SL: 26.1 mm, MD: 15.9 mm, Maria Dreieichen ( Austria), illustrated in Hörnes (1853: pl. 24, fig. 12), Figs 28G View FIGURE 28 1 –G View FIGURE 1 2 View FIGURE 2 . Paralectotypes: NHMW 1852 View Materials /0012/0009b, SL: 25.2 mm, MD: 14.3 mm, Maria Dreieichen ( Austria), Figs 28F View FIGURE 28 1 –F View FIGURE 1 2 View FIGURE 2 . NHMW 1852 View Materials /0012/0009c, SL: 25.3 mm, MD: 14.7 mm, Maria Dreieichen ( Austria), Fig. 28H View FIGURE 28 .

Additional material. 6 spec., NHMW 1870 View Materials /0054/0028, Mold ( Austria) .

Revised description. Medium-sized, moderately slender fusiform shell with gradate spire; apical angle ~53– 57°. Protoconch unknown. Teleoconch of up to five whorls. Suture linear, moderately incised. Early teleoconch whorls with broad subsutural ramp, angled shoulder, convex below. About ten prominent axial ribs, fading on subsutural ramp, overrun on antepenultimate and penultimate whorls by distinct adis, IP, abis and prominent P1, P2; s1, s2 delicate, P3 appears at abapical suture on penultimate whorl. Axial ribs and spiral cords forming cancellate sculpture. Last whorl attaining ~75% of total height, with shallow, weakly concave and slightly allometrically widening subsutural ramp, rounded shoulder, moderately convex periphery, moderately constricted below. Up to 13 weak axial ribs fading over subsutural ramp. Cords adis, IP, abis overrun by scabrose growth lines. P1–P6, P6 on convex part of base, ADP, MP, ABP prominent; P1 and P2 swollen over axial ribs, forming indistinct tubercles, P5 slightly more prominent, delimiting base; s1–s4 weaker, slightly recessed, s5, s6 slightly more prominent, ads weak. Fasciole moderately swollen, delimiting moderately narrow pseudoumbilicus. Aperture moderately wide; outer lip thickened with subosolete ID and prominent D1–D6. Anal canal indistinct, wide; siphonal canal long, open or fused, straight, weakly dorsally recurved. Columella straight, weakly twisted at transition to siphonal canal. Columellar callus forming broad, adherent rim.

Discussion. Ocenebra schoenni (Hörnes, 1853) is characterized by its peculiar cancellate sculpture and cannot be confused with any other Paratethyan species. Ocinebrina houarti Landau, Merle, Ceulemans & Van Dingenen, 2019 , from the Tortonian of France, has also a markedly cancellate sculpture but lacks secondary cords and is more globose. The extant Mediterranean Ocinebrina cyclopus Monterosato, 1884 has cancellate sculpture but also lacks distinct secondary cords and has a more incised suture.

Paleoenvironment. Typical in brackish, lagoonal deposits of the Eggenburgian Loibersdorf Formation as described by Mandic et al. (2004).

Distribution in Central Paratethys. Egerian (Late Oligocene/Early Miocene): Hungarian Paleogene Basin: Diósjenő ( Hungary) ( Báldi et al. 1964). Eggenburgian (Early Miocene): North Alpine-Carpathian Foreland Basin: Mold, Maria Dreieichen ( Austria) ( Steininger et al. 1971).

Genus Ocinebrinopsis nov. gen.

Etymology. From Ocinebrina derived from Ancient Greek ὄψις (ópsis, “aspect”, “appearance”). Gendre female.

Type species. Ocenebra orientalis Friedberg, 1928 , Middle Miocene , Central Paratethys Sea .

Diagnosis. Medium-sized biconic with conical spire with strongly developed primary cords. Typically, with IP, P1 to P6, ADP, MP and ABP, occasionally EAB1 on siphonal canal; secondary cords (adis, abis, s1 to s6) usually weaker. Subsutural collar present. Axial ribs moderately prominent. Siphonal canal short open or sealed. Internal denticles ID, D1 to D6 usually elongate or bifid.

Description. Medium-sized shell. Biconic with conical spire. Subcarinate teleoconch whorls. Shoulder placed at two-third whorl height with prominent P1 on last whorl. Broad, weakly concave subsutural ramp. Strongly developed primary cords. On spire: IP, P1–P2 or IP, P1 (convex part of whorl). On last whorl: IP (subsutural ramp), P1 to P6 (convex part of whorl), ADP, MP and ABP, occasionally EAB1 (siphonal canal); secondary cords (adis, abis, s1 to s6) usually weaker. Subsutural collar present, more or less swollen. Moderately prominent axial ribs; 8–10 on last whorl. Aperture moderately small, ovate; anal canal weakly incised; no parietal callus; columellar lip smooth, adherent; siphonal canal short open or sealed. Internal denticles ID, D1 to D6 usually elongate or bifid. Aragonitic microstructure.

Included species. Middle Miocene of Paratethys: Murex ( Occenebra [sic]) credneri (Hoernes & Auinger, 1885) ; Ocinebrinopsis gregaria sp. nov.; Ocenebra orientalis Friedberg, 1928 ; Ocinebrinopsis aperta sp. nov.; Ocinebrinopsis dominicii sp. nov.; Ocinebrinopsis aperta sp. nov.; Murex crassilabiatus Hilbert, 1879 ; Murex ( Occenebra [sic]) sublavatus Basterot var. grundensis Hoernes & Auinger, 1885 ; Tritonium striatum Eichwald, 1830 ; Ocinebrina bertai Kovács, 2020 ; Pollia weinsteigensis Hoernes & Auinger, 1890 . Middle Miocene of France: Fusus coelatus (Dujardini, 1837) . Late Miocene of Mediterranean Sea: Murex dertonensis Bellardi, 1873 .

Distribution: Early to Late Miocene (Burdigalian to Tortonian): Paratethys Sea and, Proto-Mediterranean Sea.

Discussion. The fossil species included herein in this new genus have often been attributed to the genus Ocinebrina Jousseaume, 1880 (type species: Murex corallinus Scacchi, 1836 ) due to their medium-sized shell, their biconic profile with conical spire, their short siphonal canal, sometimes sealed, and their strongly developed primary cords. Barco et al. (2017) revised present-day Ocinebrina . The shells of present day Ocinebrina differ from those of Ocinebrinopsis by having a slenderer shape, a more convex subsutural ramp and by lacking an anal notch and subsutural collar. The aperture of Ocinebrina displays usually rounded denticles (ID, D1–D6), whereas that of Ocinebrinopsis usually has elongated denticles. Paciocinebrina Houart et al. 2019 (type species: Tritonium luridum Middendorff, 1848 ) includes northeast Pacific species that often resemble Ocinebrina . However, they differ from Ocinebrina and Ocinebrinopsis by the morphology of their outer lip denticles, which comprise D1–D5 (rarely D6) and not ID, D1–D6 ( Houart et al., 2019, figs 8–10). According to Crocetta et al. (2020), shells of Ocenebra Gray, 1847 show a great intrageneric variability and some species are similar to those of Ocinebrinopsis . However, Ocinebrinopsis differs by having a concave subsutural ramp, a subsutural collar, and an anal notch. Moreover, the type species of the genus, O. erinaceus ( Linnaeus, 1758) , is trivaricate and has a calcitic layer, missing in Ocinebrinopsis .