Ocinebrinopsis orientalis (Friedberg, 1928)

Ocinebrinopsis orientalis (Friedberg, 1928)

Figs 29 View FIGURE 29 , 32A–E View FIGURE 32

Murex sublavatus Basterot View in CoL —Hörnes 1853: 236 (pars), pl. 24, fig. 15 [non Ocenebra sublavata (de Basterot, 1825) View in CoL ]. Murex View in CoL ( Occenebra [sic]) caelatus Grat. var badensis —Hoernes & Auinger 1885: 217 (pars), pl. 26, figs 13–14 [non Murex View in CoL badensis Nyst, 1881 View in CoL ]. Ocenebra sublavata Bast. View in CoL — Friedberg 1912: 177, pl. 11, figs 8–9 [non Ocenebra sublavata (de Basterot, 1825) View in CoL ; non fig. 10 =

Ocinebrinopsis sp. ]. * Ocenebra orientalis Friedb. —Friedberg 1928: 587 [pro Ocenebra sublavata sensu Friedberg 1912 View in CoL , pl. 11, figs 8–10]. Ocenebra orientalis Friedb. — Friedberg 1938: 116. Murex aff. craticulatus View in CoL L.— Strausz 1943: 144, pl. 3, figs 7–8. Tritonalia sublavata Bast. — Strausz 1954: 25 (pars), pl. 4, figs 95a–b [non Ocenebra sublavata (de Basterot, 1925) View in CoL ]. Ocinebrina sublavata dertonensis Mayer View in CoL (in Bellardi), 1872— Strausz 1966: 273, pl. 33, figs 7–8 [non Ocinebrinopsis dertonensis

( Bellardi, 1873)]. Tritonalia (Ocinebrina) orientalis (Friedberg 1928) — Kojumdgieva in Kojumdgieva & Strachimirov 1960: 150, pl. 40, fig. 10. Tritonalia orientalis ( Friedberg, 1912) — Zelinskaya et al. 1968: 188, pl. 44, figs 19–20. [ Ocinebrina View in CoL ] badensis Hoernes & Auinger 1885 View in CoL — Vokes 1971: 21. Ocinebrina sublavatus dertonensis (Mayer) View in CoL — Stancu et al. 1971: 125, pl. 8, fig. 6 [non Ocinebrinopsis dertonensis ( Bellardi,

1873)]. Tritonalia sublavata Bast. — Csepreghy-Meznerics 1972: 27, pl. 9, figs 23–24 [non Ocenebra sublavata (de Basterot, 1825) ]. Tritonalia sublavata dertonensis May.— Csepreghy-Meznerics 1972: 27, pl. 9, figs 27–28 [non Ocinebrinopsis dertonensis

( Bellardi, 1873)]. Tritonalia caelata Grat. — Csepreghy-Meznerics 1972: 27, pl. 10, figs 1–2 [non Ocinebrinopsis coelata ( Dujardin, 1837) ]. Tritonalia caelata (Grateloup) 1833 — Bohn-Havas 1973: 1113: pl. 5, figs 12–13 [non Ocinebrinopsis coelata ( Dujardin,

1837)].? Ocinebrina coelata badensis (R. Hoernes et M. Auinger, 1885) — Švagrovský 1982: 387, pl. 1, fig. 5. Ocinebrina coelata badensis (Hoernes, Auinger) — Nikolov 1994: 51, pl. 1, figs 9–11, pl. 2, figs 3–6. Hadriania credneri (Hoernes & Auinger, 1885) — Bałuk 1995: 225, pl. 28, fig. 8 [non Ocinebrinopsis credneri (Hoernes &

Auinger, 1885)]. Ocinebrina credneri (Hoernes et Auinger) — Kovács & Vicián 2024: 24, figs 59–61 [non Ocinebrinopsis credneri (Hoernes &

Auinger, 1885)].

Type material. Lectotype (designated herein) of Murex orientalis Friedberg, 1912 , State Natural History Museum, National Academy of Sciences of Ukraine, L’viv); we have not been able to receive a picture of the specimen; SL: 24 mm, MD: 13 mm, Korytnica ( Poland), illustrated in Friedberg (1912: pl. 11, fig. 8).

Lectotype of Ocinebrina badensis (Hoernes &Auinger, 1885) designated by Švagrovský (1982: 387), NHMW 1869/0001/0079a, SL: 23.2 mm, MD: 11.1 mm, Baden-Sooss ( Austria), illustrated in Hoernes & Auinger (1885: pl. 26, fig. 13) (the same specimen was designated as lectotype by Nikolov 1994: 51), Figs 32A View FIGURE 32 1 –A View FIGURE 1 3 View FIGURE 3 . Paralectotype of Ocinebrina badensis (Hoernes &Auinger, 1885) : NHMW 1869/0001/0079b, SL: 18.2 mm, MD: 8.6 mm, Baden-Sooss ( Austria), illustrated in Hoernes & Auinger (1885: pl. 26, fig. 14), Figs 32B View FIGURE 32 1 –B View FIGURE 1 3 View FIGURE 3 .

Illustrated material. NHMW 1869/0001/0035, SL: 25.8 mm, MD: 13.8 mm, Bad Vöslau, illustrated in Hörnes (1853: pl. 24, fig. 15), Figs 32C View FIGURE 32 1 –C View FIGURE 1 3 View FIGURE 3 . NHMW 2024/0047/0002, SL: 24.8 mm, MD: 12.4 mm, Bad Vöslau ( Austria), Figs 32D View FIGURE 32 1 –D View FIGURE 1 3 View FIGURE 3 . NHMW 2024/0047/0002a, SL: 24.6 mm, MD: 13.3 mm, Bad Vöslau ( Austria), Figs 32E View FIGURE 32 1 –E View FIGURE 1 2 View FIGURE 2 .

Revised description. Medium-sized, slender fusiform shell with high spire; apical angle ~55°. Protoconch unknown. Teleoconch of up to six whorls; suture weakly incised, shallowly undulating. Early teleoconch whorls weakly shouldered with about nine broad axial ribs with rapidly widening interspaces, fading over subsutural ramp. Broad, weakly concave subsutural ramp with weakly scabrose cord, adis, IP, abis distinct (on penultimate and antepenultimate whorls) with slightly wider interspace between adis and IP; tertiary threads may be present. Shoulder placed mid-whorl with prominent P1, swollen over axial ribs; P2 prominent, s1, s2 weaker. Last whorl attaining ~73–77% of total height; with concave subsutural ramp, angled or slightly rounded shoulder, weakly convex to subcylindrical periphery, moderately constricted below. Spiral sculpture of broad, scabrose subsutural collar, prominent adis, IP, abis. P1 most prominent, forming blunt tubercles over axial ribs. P1–P6 (P6 on convex part of base), ADP, MP, ABP, EAB1, s4–s5 occasionally prominent, convex of subequal strength; s1–s3 slightly weaker. One narrow tertiary thread in interspaces between primary and secondary cords at periphery. Fasciole strongly swollen, twisted, delimiting distinct, moderately wide pseudoumbilicus. Aperture narrowly ovate; outer lip thickened by terminal varix, with crenulate edge, bearing ID, D6 making short lirae close behind peristome; ID simple, D1–D6 occasionally bifid. Anal canal moderately incised, poorly delimited. Siphonal canal moderately long, narrow, fused. Slightly bent to the left, weakly dorsally recurved at tip. Columella weakly excavated, smooth, slightly twisted at siphonal canal. Columellar callus forming broad rim, sharply delimited, weakly erect at siphonal canal, adherent in parietal area.

Paratethyan synonyms. When introducing Murex ( Occenebra [sic]) caelatus badensis as the name for this species, Hoernes & Auinger (1885) overlooked that Murex badensis was already preoccupied by Nyst (1881). When establishing Murex condigus, De Gregorio (1885a: 245) also discussed Murex sublavatus of Hörnes 1856: 236, pl. 24, figs 14–15] but Murex condigus is based on syntypes from Ciminna ( Italy), and therefore it is not a synonym of Ocinebrinopsis orientalis .

Discussion. This species is characterized by its slender outline and fused siphonal canal (note that the canal is damaged in Figs 32A, E View FIGURE 32 ). It differs from Ocinebrinopsis credneri (Hoernes & Auinger, 1885) in its smaller size, higher spire and narrower subsutural ramp. Ocinebrinopsis dominicii sp. nov. is slenderer and has a longer and open siphonal canal.

Paleoenvironment. Occurrences in the Baden Formation of the Vienna Basin suggest middle to outer neritic environments in up to 250 m water depth ( Kranner et al. 2021) .

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica( Poland) (Friedberg 1928); Vienna Basin: Baden-Sooss, Bad Vöslau ( Austria) (Hoernes & Auinger 1885); Borský Mikuláš ( Slovakia) ( Švagrovský 1982); Pannonian Basin: Márkháza, Várpalota ( Hungary) ( Kovács & Vicián 2024; Strausz 1966); Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972); Southern Carpathians: between Dubova and Pojejena ( Romania) ( Stancu et al. 1971); Făget Basin: Coşteiu de Sus ( Romania) (Hoernes & Auinger 1885); Dacian Basin: Dobrusha, Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960; Nikolov 1994).