Papuanatula, Kluge & Novikova, 2011

Kaltenbach, Thomas, Kluge, Nikita J. & Gattolliat, Jean-Luc, 2025, Redescription of Papuanatula Lugo-Ortiz & McCafferty (Ephemeroptera, Baetidae), with description of a new subgenus and 20 new species, ZooKeys 1227, pp. 159-347 : 159-347

publication ID	https://doi.org/10.3897/zookeys.1227.138100
publication LSID	lsid:zoobank.org:pub:F259B26F-4DA1-452E-ABEE-7D0957CFE261
DOI	https://doi.org/10.5281/zenodo.14853758
persistent identifier	https://treatment.plazi.org/id/F7F36491-7598-5A59-BCB1-B831EB479A93
treatment provided by	ZooKeys by Pensoft
scientific name	Papuanatula
status	s. str.

Treatment

Subgenus Papuanatula s. str.

Type species.

Papuanatula (Papuanatula) tuber Lugo-Ortiz & McCafferty, 1999 .

Diagnosis (larval characters).

Distal part of antenna asymmetrical: each flagellomere with anterior side more convex than posterior side, with brown hypodermal spot near apex of anterior side (Fig. 2 b View Figure 2 ). Labrum dorsally with submarginal row of long, feathered setae (Fig. 3 b View Figure 3 ). Each seta consists of a stout stem and long processes on both sides. Mandibles. Both mandibles long, with incisor strongly elongated (in contrast to shortened in Papuafiliola nov.); full-length mandibles present only at the beginning of each instar, often worn at the end of instar. Right mandible (Fig. 3 e View Figure 3 ): kinetodontium deeply separated from incisor; incisor with three denticles; kinetodontium terminating with three or four denticles, with distal denticle longest. Left mandible (Fig. 3 f View Figure 3 ): incisor and kinetodontium fused at most of its length; incisor with three denticles; kinetodontium terminating with three or four denticles, with distal denticle longest. Labium: glossa usually approximately as long as half of paraglossa; with several long setae at apex and usually one long seta near middle of ventral side. Labial palp without real distomedian projection on segment II (Fig. 3 c View Figure 3 ) (in contrast to Papuafiliola subgen. nov.). Legs (Figs 4 a – e View Figure 4 , 108 b View Figure 108 ): Outer side of each femur usually with single regular row of long, hair-like setae bearing numerous fine, short branches on all sides. Anterior side of each tibia usually with regular row of setae similar to that on femur. Anterior side of each tarsus usually with regular row of setae similar to femur and tibia. Posterior side of each tarsus with regular row of short, stout setae and one (rarely two) much longer, thinner, pointed seta distad. Paraproct (Fig. 12 e View Figure 12 ): posterior margin usually with prolongation. Pose of subimaginal gonostyli under larval cuticle (Fig. 76 b View Figure 76 ). Subimaginal gonostyli developing under cuticle of last instar male larvae folded in “ Labiobaetis - type ” ( Kluge 2004: fig. 29 I): 1 st segment directed laterally, 2 nd and 3 rd segments directed medially, whereby the 2 nd segment is the closest to the posterior margin of the 9 th segment.