Hypselostoma hungerfordianum Möllendorff, 1891

Hypselostoma hungerfordianum Möllendorff, 1891

Figs 186 F View Figure 186 , 196 View Figure 196 , 197 View Figure 197 , 198 View Figure 198 , 199 View Figure 199 , 200 View Figure 200 , 201 View Figure 201

Hypselostoma hungerfordianum Möllendorff, 1891: 337, pl. 30, figs 7, 7 a. View in CoL

Hypselostoma striolatum Möllendorff, 1894: 152. syn. nov.

Hypselostoma hungerfordianum View in CoL — Sykes 1902: 61.

Hypselostoma striolatum — Fischer and Dautzenberg 1904: 408.

Gyliauchen hungerfordianus — Pilsbry 1917: 212, pl. 36, figs 1–4.

Gyliauchen striolatus — Pilsbry 1917: 215.

Gliotrachela hungerfordiana [sic] — Laidlaw 1933: 214.

Gliotrachela striolatus [sic] — Laidlaw 1933: 214.

Gyliotrachela hungerfordiana View in CoL — van Benthem Jutting 1949 b: 60; van Benthem Jutting 1950: 46; van Benthem Jutting 1950: 26; Zilch 1959: 164, fig. 563; van Benthem Jutting 1960: 14; Zilch 1984: 166; Davison 1995: 239; Schileyko 1998: 142, fig. 162; Maassen 1999: 122–123; Schilthuizen 1999: 283; Maassen 2001: 75–76; Foon et al. 2017: 79, fig. 30 B; Hoekstra and Schilthuizen 2011: 267–272, fig. 2; Foon and Marzuki 2023: 142, fig. 1 AP; Tongkerd et al. 2024: 180–182 View Cited Treatment , figs 10, 13 M.

Gyliotrachela striolata View in CoL — van Benthem Jutting 1950: 46; Zilch 1984: 166; Thompson and Upatham 1997: 233; Hemmen and Hemmen 2001: 41.

Gyliotrachela khaochongensis Panha in Panha 1998 a: 53 –56, fig. 2. View in CoL

Gyliotrachela khaochongensis View in CoL — Hemmen and Hemmen 2001: 41; Panha and Burch 2008: 70–71, fig. 61. Tongkerd et al. 2024: 180, fig. 10 B.

Hypselostoma satulensis Panha & Burch in Panha et al. 2004: 75–76, fig. 12. syn. nov. View in CoL

Hypselostoma satulensis View in CoL — Panha and Burch 2008: 98–99, fig. 84; Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.

Gyliotrachela striolatus View in CoL [sic] — Panha and Burch 2008: 81–82, fig. 71.

Gyliotrachela phoca Tongkerd & Panha, 2013 in Tongkerd et al. 2013: 71–75, figs 5–7. View in CoL

Gyliotrachela phoca View in CoL — Tongkerd et al. 2024: 180. Dumrongrojwattana and Wongkamhaeng 2024: 324, fig. 8.

Type material examined.

Malaysia • lectotype of H. hungerfordianum ; SMF 4587. Thailand • holotype of H. striolatum ; SMF 4591 • holotype of H. satulensis ; S. Panha leg.; CUMZ ver. 44028; paratype of G. khaochongensis ; from the type locality; CUMZ ver. 010.

Additional material examined.

Thailand-South • 6 shells; Suratthani Province, 6 km S of Na San , limestone mountain; 8.660°N, 99.398°E; 03 June 1987; F. Thompson leg.; locality code FGT-4311 ; UF 530647 GoogleMaps • 2 shells; Suratthani Province, Hwy. 401, 2.7 km W junction Hwy . 4142 and 401, evergreen forest on rocky hillside; 9°10'N, 99°40'E; 90 m a. s. l.; 19 Apr. 1988; K. Auffenberg leg.; locality code KA-0674 ; UF 591325 GoogleMaps • 34 shells; Suratthani Province, 15.3 km NE Ban Na San off road to Tha Rom Yen National Park ; 8°53'N, 99°26'E; 40 m a. s. l.; 20 Apr. 1988; K. Auffenberg leg.; locality code KA-0678 ; UF 345386 GoogleMaps • 12 shells; Suratthani Province, 1 km NE of Na San , 100 m limestone mountain; 8.812°N, 99.382°E; 02 June 1987; F. G. Thompson leg.; locality code FGT-4309 ; UF 547905 GoogleMaps • 3 shells; Suratthani Province, Angthong islands, Wia Talap island, steep path to Buaboke cave , at the base of limestone rocks; 8°7.603'N, 98°55.466'E; 50 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI GoogleMaps • 1 shell; Suratthani Province, 15.3 km NE Ban Na San off road to Tha Rom Yen National Park ; 8°53'N, 99°26'E; 40 m a. s. l.; 20 Apr. 1988; K. Auffenberg leg.; locality code KA-0678 ; UF 591326 GoogleMaps • 3 shells; Phangnga Province, 32 km NE Khok Kloi, Wat Tham Suwan Khuha ; 08°27'N, 98°27'E; 20 m a. s. l.; 27 Mar. 1988; K. Auffenberg leg.; locality code KA-0601 ; UF 345751 GoogleMaps • 1 shell; Phangnga Province, limestone knoll, 5.7 km SW intersection Hwys 4 & 4152; 8°27'N, 98°29'E; 20 m a. s. l.; 28 Mar. 1988; K. Auffenberg leg.; locality code KA-0602 ; UF 345771 GoogleMaps • 1 shell; Krabi Province, 5.6 km SW of Ao Luek Tai , on uneven surface of large boulder at the base of limestone cliff; 8°20.772'N, 98°40.968'E; 20–25 m a. s. l.; 16 Mar. 2020; ex. coll. S. Aiken; coll. HA GoogleMaps • 8 shells; Krabi Province, 9.2 km northwest from Phang Nga ; 08°07.305'N, 98°52.231'E; 40 m a. s. l.; 21 Feb. 2015; A. Hunyadi leg.; coll. HA GoogleMaps • 3 shells; Krabi Province, Viewpoint Hill, Railay (= Rai Leh ) Beach West , at the base of limestone rocks; 8°0.511'N, 98°50.248'E; 40 m a. s. l.; Sep. 2007; A. Reischütz leg.; coll. REI GoogleMaps • 4 shells; Nakhon Si Thammarat Province, Lan Saka, N side of Hwy . 4015; 08°25'N, 99°45'E; 16 Apr. 1988; K. Auffenberg leg.; locality code KA-0665 ; UF 345226 GoogleMaps • 8 shells; Nakhon Si Thammarat Province, circa 38 km WNW of Tha Sala, outcrop 1.5 km W of junction Hwys 4186 and 4188; 8°43'N, 99°47'E; 80 m a. s. l.; 16 Apr. 1988; K. Auffenberg leg.; locality code KA-0661 ; UF 345194 GoogleMaps • 16 shells; Nakhon Si Thammarat Province, Wat Tham Thong Panara , vicinity of the rock temple; 08°25.278'N, 99°22.762'E; 19 Feb. 2015; A. Hunyadi leg.; coll. HA GoogleMaps • 23 shells; Nakhon Si Thammarat Province, circa 38 km WNW of Tha Sala, outcrop 1.5 km W of junction Hwys 4186 and 4188; 08°43'N, 99°47'E; 80 m a. s. l.; 16 Apr. 1998; K. Auffenberg leg.; locality code KA-0661 ; UF 345193 GoogleMaps • 20 shells; Phattalung Province, 6 km NW of Phattalung, 1 km NE of Hwy. 4048; 7°40'N, 100°3'E; 60 m a. s. l.; 13 Apr. 1988; K. Auffenberg leg.; locality code KA-0653 ; UF 345102 GoogleMaps • 54 shells; Phatthalung Province, Khao Pu-Khao Ya Nat. Park, Si Banphot district, Ruesi Cave ; 7°41.797'N, 99°50.162'E; 25 Feb. 2023; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. PGB GoogleMaps • 25 shells; Phattalung Province, 6.4 km E of Hwy. 4 on Hwy. 4081, 1 km SW of Khao Chai Son ; 7°27'N, 100°11'E; 50 m a. s. l.; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0648 ; UF 345063 GoogleMaps • 14 shells; Phattalung Province, 6.4 km E of Hwy. 4 on Hwy. 4081, 1 km SW of Khao Chai Son ; 7°27'N, 100°11E; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0648 ; UF 345064 GoogleMaps • 7 shells; Phattalung Province, 16.2 km S of junction of Hwy . s 4 and 4122; 7°25'N, 99°59'E; 150 m a. s. l.; 09 Apr. 1988; K. Auffenberg leg.; locality code KA-0637 ; UF 344971 GoogleMaps • 4 shells; Phattalung Province, 6 km NW of Phattalung, 1 km NE of Hwy. 404; 7°40.02'N, 100°3'E; 60 m a. s. l.; 13 Apr. 1988; K. Auffenberg leg.; locality code KA-0653 ; UF 436278 GoogleMaps • 18 shells; Phattalung Province, 15.1 km W Hwy. 41 on Hwy. 4164, Khaopu and Khaoya National Park ; 7°47'N, 99°56'E; 160 m a. s. l.; 14 Apr. 1988; K. Auffenberg leg.; locality code KA-0657 ; UF 345154 GoogleMaps • 6 shells; Phattalung Province, 6.4 km E Hwy. 4 on Hwy. 4081, 1 km SW Khao Chai Son ; 7°27'N, 100°10.98'E; 11 Apr. 1988; K. Auffenberg leg.; locality code KA-0649 ; UF 345073 GoogleMaps • 17 shells; Phattalung Province, 16.2 km S of junction of Hwys 4 and 4122; 7°25'N, 99°59'E; 150 m a. s. l.; 09 Apr. 1988; K. Auffenberg leg.; locality code KA-0637 ; UF 344970 GoogleMaps • 3 shells; Phattalung Province, Khao Pu Khao Ya National Park, vicinity of Matcha cave ; 7°39.980'N, 99°52.343'E; 125 m a. s. l.; 25 Feb. 2023; A. Hunyadi leg.; coll. HA GoogleMaps • 23 shells; Phattalung Province, Khao Pu Khao Ya National Park, Si Banphot district, ca 6 km north-northwest from Taphaen ; 7°40.738'N, 99°50.822'E; 85 m a. s. l.; 24 Feb. 2023; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA GoogleMaps • 1 shell; Yala Province, 6.3 km W of Yala, Ban Na Tham, Wat Khuha Phi Muk ; 6°33'N, 101°15'E; 40 m a. s. l.; 01 Apr. 1988; K. Auffenberg leg.; locality code KA-0612 ; UF 344665 GoogleMaps • 3 shells; Yala Province, 6.3 km W of Yala, Ban Na Tham, Wat Khuha Phi Muk ; 6°33'N, 101°15'E; 40 m a. s. l.; 01 Apr. 1988; K. Auffenberg leg.; locality code KA-0612 ; UF 344664 GoogleMaps • 25 shells; Yala Province, Mueang Yala district, Na Tham, Silp Cave (~ 9 km from centre of Yala ); 6°31.316'N. 101°13.982'E; 21 Feb. 2023; A. Hunyadi, K. Okubo & J. U. Otani leg.; coll. HA. Malaysia GoogleMaps • 2 shells; Selangor, Templer Park , Bukit Takun; NNW of Kuala Lumpur; Nov. 1998; ex. coll. Hemmen; coll. PGB • 46 shells; Kedah, Baling, eastern side of Bukit Baling ; 55 m a. s. l.; 05°40.575'N, 100°54.803'E; 11 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 9 shells; Kelantan, Gua Musang , environment of the cave entrance; 04°52.974'N, 101°58.116'E; 135 m a. s. l.; 17 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 7 shells; Perlis, Kaki Bukit, vicinity of Gua Kelam ; 06°38.674'N, 100°12.170'E; 45–55 m a. s. l.; 13 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 11 shells; Pahang, 15 km west from Bandar Pusat Jengka, Hutan Lipur Gunung Senyum ; 03°41.862'N, 102°25.980'E; 85 m a. s. l.; 23 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 3 shells; Perak, 800 m north from Padang Rengas on the road no. A 107, eastern side of Gunung Pondok ; 04°47.195'N, 100°50.493'E; 90 m a. s. l.; 08 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 12 shells; Kelantan, Gua Musang, 6 km Pulai, Gua Madu , vicinity of the cave; 04°50.213'N, 101°56.982'E; 120 m a. s. l.; 17 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 3 shells; Selangor, Kuala Lumpur E - Kanching, Templer Park, Perangsang Templer Golf Club , rock; 03°17.878'N, 101°38.438'E; 100 m a. s. l.; 25 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 22 shells; Perak, Lenggong, Taman Awam Gua Tok Giring ; 05°07.128'N, 100°.351'E; 90 m a. s. l.; 06 Jan. 2013; A. Hunyadi leg.; coll. HA • 131 shells; Perak, northeast from Chemor, 2 km east from Kampung Kanthan , western side of the mountain; 04°44.799'N, 101°07.794'E; 105 m a. s. l.; 08 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 304 shells; Pahang, Bukit Cinta Manis, south-southeastern side, 800 m from Lebuhraya Karak towards Kampung Cinta Manis ; 03°26.714'N, 102°00.814'E; 130 m a. s. l.; 22 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 18 shells; Perlis, Kangar, Bukit Kaya / Bukit Lagi , northeastern edge of the mountain; 06°26.076'N, 100°11.469'E; 5 m a. s. l.; 12 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 67 shells; Pahang, Gua Bama, Kuala Lipis 9 km, Padang Tungku ; 04°11.652'N, 101°57.936'E; 120 m a. s. l.; 19 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 180 shells; Pahang, 20 km southeast from Jerantut, Gua Kota Gelanggi, below Gua Balai ; 03°54.000'N, 102°28.412'E; 115 m a. s. l.; 21 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 4 shells; Kelantan Province, Gua Musang , behind railway station; 04°53.033'N, 101°58.178'E; J. U. Otani leg.; coll. PGB GoogleMaps • 6 shells; Pahang Province, Lipis, Kenong Rimba park; 04°12.952'N, 102°11.263'E; 83 m a. s. l.; J. U. Otani leg.; coll. PGB GoogleMaps • 2 shells; Pahang Province, Jerantut, Taman Negara, Kuala Tahan , Gua Luas ; 04°30.640'N, 102°26.548'E; 75 m a. s. l.; J. U. Otani leg.; coll. PGB GoogleMaps • 1 shell; Pahang Province, Jerantut, Kg. Baheru, Gua Kota Gelanggi , Gua Terang Bulan; J. U. Otani leg.; coll. PGB • 8 shells; Kedah, 1.5 km east from Kodiang, Gua Kerbau ; 06°23.431'N, 100°18.904'E, 14 Jan. 2013; A. Hunyadi leg.; coll. HA GoogleMaps • 5 shells; Kelantan Province, Gua Musang, 264 km from KL, Route 8; 191 m a. s. l.; 04°46.685'N, 102°00.095'E; J. U. Otani leg.; coll. PGB GoogleMaps • 5 shells; Selangor, Templer Park , Bukit Takun; NNW of Kuala Lumpur; Nov. 1999; J. Hemmen leg.; coll. PGB • 5 shells; Kelantan Province, Gua Madu, south of Gua Musang ; 04°52.405'N, 101°57.925'E; 02 Nov. 2000; J. Hemmen leg.; coll. PGB GoogleMaps • 8 shells; Pahang, Kota Gelanggi, Jengka-Jerantut road, on limestone surfaces covered with mosses; 03°53.4758'N, 102°28.388'E; 29. May 2011; M. E. Marzuki leg.; coll. PGB GoogleMaps .

Type localities.

“ Perak ” (Bukit Pondong, Malaysia) ( H. hungerfordianum ); “ Kuankalong Limestone Hill, Satul Province, 6°52'59"N, 100°07'38"E, 110 meters elevation ”, Thailand ( H. satulensis ); “ Samui islands, Gulf of Siam ”, Thailand ( H. striolatum ).

Differential diagnosis.

See under H. sichang and H. loei .

Distribution.

This species is widely spread in Malaysia and adjacent regions of Thailand. Also known from Myanmar where the northernmost locality of the species is known ( Tongkerd et al. 2024). The southernmost localities are positioned in and near the city of Kuala Lumpur ( Malaysia).

Remarks.

This species can have a more or less elongated spire. The umbilicus is usually consistent in its width but we have observed slight variations. The apertural dentition is variable in terms of the number of smaller barriers between the five main ones (angular, parietal, upper palatal, lower palatal and columellar). Gyliotrachela striolata was known only from a single damaged type, which retained its last whorl only. Even in damaged condition, it was clear that there are no differences between G. striolata and H. hungerfordianum which is why they are treated conspecific. Hypselostoma satulensis was described from Satun province in S Thailand and shows no morphological differences from H. hungerfordianum . Its type locality is practically located in the middle of the distribution area of H. hungerfordianum . We have examined the material from northern Thailand hosted in UF with the following data: 2 shells; Mae Hong Son Province, Tham Lod Cave, 1.5 km SE Ban Tham Lod Village; 19°34'N, 98°9'E; 720 m a. s. l.; 19 Mar. 1988; K. Auffenberg leg.; locality code KA-0576 ; UF 345527. Since this locality is quite far from the actual distribution area of this species, we believe that this sample was mislabelled and that this species actually does not occur in Mae Hong Son Province.