Agrypon laevifrontator PÉNIGOT, 2025

Agrypon laevifrontator PÉNIGOT nov.sp. ( Figs 4A, B, C View Fig )

Holotype: 1♀ "SW GEORGIA Kintrishi NP. 1264 m asl 41.44388824°N 41.0502904°E MT4 . 5- 20.V.2018 leg. G. Japoshvili " // " Anomaloninae ♀ det. Riedel 2021 " // " Holotypus Agrypon laevifrontator PÉNIGOT 2024 ". GoogleMaps

Paratypes: 1♀ "SW GEORGIA Kintrishi NP. 1634 m asl 41.44535308°N 41.05384144°E MT4 . 19.V.- 1.VI.2018 leg. G. Japoshvili " // " Paratypus Agrypon laevifrontator PÉNIGOT 2024 " GoogleMaps ; and 1♀ "SW GEORGIA Kintrishi NP. 1634 m asl 41.44535308°N 41.05384144°E MT12 . 5-20.V.2018 leg. G. Japoshvili " // " Paratypus Agrypon laevifrontator PÉNIGOT 2024 " GoogleMaps .

D e s c r i p t i o n o f t h e H o l o t y p e:

Head. Distance between the base of the antennae and the apex of the clypeus 1.14 × longer than the minimal width of the face. Eyes weakly converging ventrally, maximal width of face 1.38 × longer than its minimal width. Face centrally finely punctate, superficially rugulose, and weakly shining, the lateral margins of the face smooth and shining. Clypeus 1.81 × wider than high, smooth, shining, with some superficial punctures on its upper part. Apical margin of clypeus raised with a distinct median tooth. Malar space 0.3 × as long as the mandibular base. Lower mandibular tooth 0.45 × as long as the upper. Frons smooth, only superficially rugulose-punctate above the antennae, with a low but distinct vertical median carina. Lateral and upper parts of frons smooth with very thin, almost indistinct, punctures. Temples quite short and convexly narrowed behind the eyes, 0.58 × as long as the width of the eye in lateral view. Temples smooth and shining, with very thin and sparse punctures posteriorly, the distance between two points about 4 × longer than the diameter of the punctures. Occipital carina complete, joining the hypostomal carina slightly after the mandibular base. OOD 1.7 ×, and IOD 1.2 × longer than the maximal diameter of the lateral ocellus. Antennae with 38 flagellomeres. 1 st flagellomere 5.25 × longer than wide and 1.6 × longer than the 2 nd. 10 th flagellomere 3.0 ×, and subapical flagellomeres 2.3 × longer than wide.

Mesosoma. Antero-ventral corner of the pronotum rounded, without tooth. Pronotum irregularly longitudinally striate ventrally, but with a large smooth and shining area dorsally. Dorsal margin of pronotum longitudinally impressed, this groove with transverse carinulae. Epomia distinct, joining the anterior margin of the mesoscutum. Anterior edge of mesoscutum depressed. Notauli quite weak but distinct and complete. Mesoscutum 1.1 × longer than wide in dorsal view, smooth and shining, with moderately thin and sparse punctures, the distance between two points about as long as the diameter of the punctures. Posterior 0.25 of the median lobe of mesoscutum reticulate. Scutellum flat, reticulate, with distinct lateral carinae on the basal 0.8. Mesopleura shining, with thin punctures in the lower half, centrally with a smooth area, and with some irregular carinulae in the upper third. Speculum smooth and shining. Prepectal carina only slightly higher ventrally than laterally, and weakly bilobed behind the fore coxae. Lateral part of the prepectal carina extending beyond the mid-height of the mesopleura, fused with irregular reticulations just before the anterior margin of the mesopleura. Mesosternal suture smooth, without transverse carinulae. Postpectal carina interrupted in front of the mid coxae. Metasternal tooth small but present. Propodeum coarsely reticulate, as long as wide in dorsal view. Apex of propodeum reaching the apical 0.5 of the hind coxae.

Wings. Fwl = 5.2 mm. CI = 0.3. Discoidella absent, nervellus slightly curved. Hind wing with 6 distal hamuli.

Legs. Ventral face of the fore coxae with a distinct carina extending on the internal, anterior, and posterior edges. Hind coxa 2.7 × longer than high in lateral view. Hind trochanter 1.45 × longer than the trochantellus in ventral view. Hind femora 5.9 × longer than high in lateral view, and 0.63 × as long as the hind tibia. Hind basitarsus 0.50 × as long as the hind tibia, 2.4 × longer than the 2 nd tarsomere, and 8.8 × longer than wide. 2 nd hind tarsomere 4.6 × longer than wide. Hind tarsal claws hardly longer than the pulvillus, weakly curved, and with very short indistinct pectination basally.

Metasoma. Spiracle of 1 st metasomal segment situated on the apical 0.73. Postpetiole 1.6 × longer than wide in dorsal view. T 2 1.2 × longer than T 1, and 8.9 × longer than high in lateral view. T 3 0.45 × as long as T 2, and 1.55 × longer than high in lateral view. OBI = 0.95.

Colour. Head, mesosoma and coxae with short whitish pubescence. Head black. Face yellow. Clypeus, mandibles (except for the blackish teeth), malar space, and lower third of outer orbits, yellow. Vertex with a small (slightly larger than the lateral ocellus) yellow spot. Palpi pale yellow. Antennae dark brown dorsally, pale brown ventrally, except for the mostly black 1 st flagellomere. Scapus black, yellow beneath. Pedicellus entirely black. Mesosoma black, the upper hind corner of the pronotum and the subtegular ridge with a small yellowish spot. Tegula yellow. Wings hyalin with brown veins. Pterostigma greyishbrown. Fore and mid legs entirely pale yellow, except for the brownish 5 th tarsomere. Hind coxa, trochanter and trochantellus, black. Hind femora brownish, reddish basally and medio-ventrally, with a subbasal dark ring. Hind tibia reddish medially, reddish-brown basally and subapically, black on the apical 0.2. Hind tarsus mostly yellow, basitarsus with the 0.75 basal reddish-brown, 5 th tarsomere brown. Petiole brown, with the postpetiole red. T 2 mostly brownish-black, with only the ventral margin red. T 3-4 brownish dorsally, red on the ventral half. T 5 black with the antero-ventral corner broadly red. T 6-8 black, with the ventral margin yellow. Sheath and ovipositor reddish-yellow.

Male unknown. Intraspecific variability. Fwl = 4.7-5.2 mm. Antennae with 38-40 Flagellomeres. Temples 0.57-0.63 × as wide as the eye in lateral view. OOD 1.7-1.9 × the maximal diameter of the lateral ocellus. Hind femora 5.9-6.2 × longer than high in lateral view. OBI 0.95-1.0. Frons above antennae varying from almost entirely smooth, to superficially rugulo-punctate.

E t y m o l o g y: The name refers to the very weakly sculptured frons compared to the other western Palearctic Agrypon species of the A. varitarsum group.

B i o l o g y: Unknown.

D i a g n o s i s: This species is closely related to Agrypon varitarsum (WESMAEL, 1849) and Agrypon sp. A in PÉNIGOT, 2021 – the later should belong to A. cognatum FÖRSTER ,

1860 (PÉNIGOT in prep.) – because it shares the following characters: mesoscutum smooth and shining between the rather sparse punctures, and with distinct notauli; carinated fore coxae; 1 st flagellomere less than 2 × longer than the 2 nd; upper lateral part of pronotum and the upper half of mesopleura with a smooth unpunctate shining area; long and thin antennae, with the 2 nd flagellomere about 4.0-4.5 × longer than wide. It differs from Agrypon varitarsum and Agrypon sp. A by the typical weakly sculptured frons ( Figs 4B, C View Fig ), smaller ocelli, and black hind coxae.