Anomala fenestrata Zhao, Fujioka & Zorn, 2025

Anomala fenestrata Zhao, Fujioka & Zorn View in CoL , sp. nov.

[Chinese vernacular name: Oi叶异丽aek]

( Figs 9A–B View FIGURE 9 , 12D View FIGURE 12 , 23E–H View FIGURE 23 )

Type material ( 14♂♂, 21♀♀). Holotype: ♂ ( SCAU) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill. , 1753 m, 29.214870°N, 95.169272°E, 18–30.VII.2023 (at light), Yao-Nan Zhang leg.” GoogleMaps Paratypes: 3♂♂, 4♀♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill. , 1753 m, 29.214870°N, 95.169272°E, 18–30.VII.2023 (at light), Yao-Nan Zhang leg.” GoogleMaps ; 2♂♂, 5♀♀ ( ZYPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill. , 1753 m, 29.214870°N, 95.169272°E, 18–30.VII.2023 (at light), Yao-Nan Zhang leg.” GoogleMaps ; 1♂, 4♀♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng, Gelin Vill. , 1768 m, 29.21315309°N, 95.17111894°E, 2023.VII.13-14, Hao-Yi Liu leg.” GoogleMaps ; 1♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., 96k Marangkang vill., 1305 m, 29.56287354°N, 95.46563172°E, 2023.VII.17, Hao-Yi Liu leg.” GoogleMaps ; 1♂, 4♀♀ ( ZMPC) “ CHINA: Tibet, Nyingchi City , Medog County, 1143 m 29°19′22.62″N, 95°20′1.65″E 26.VII.2022 (at light) Chuan-Tao Qu leg.” GoogleMaps ; 1♂ ( ZMPC) “ CHINA: Tibet, Nyingchi City , Medog County, 1143 m 29°19′22.62″N, 95°20′1.65″E 20.VII.2022 (at light) Chuan-Tao Qu leg. GoogleMaps ; 1♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge , 29.62527885°N, 95.48554466°E, 1681 m, 2023.VII.9, Xu-Hong-Yi Zheng” GoogleMaps ; 2♂♂, 1♀ ( MFPC) “ China-India border: alt. 1700m Lower Subansiri 18.V. – 01.VI. 2017 Takeshi Maeda-leg.” ; 1♀ ( MFPC) “ China-India border: Lower Subansiri 9–19. AUG. 2018 Takeshi Maeda-leg.” ; 3♂♂ ( MFPC) “ China-India border: W. Siang 03. JULY. 2011 Takeshi MAEDA-leg.”

Description. Male. Body shape elongated ovoid, somewhat subparallel, moderately convex.

Color . Generally yellowish brown, the head, antennal club, disc of pronotum except for a narrow lateral area, scutellum, mesosternum, metasternum (except for a square marking laterally), pygidium in anterior half, abdominal ventrites 1–5 (except for lateral portion), meso- and metatibiae as well as all tarsomeres dark brown, protibia only blackish brown at dorsal face, protibial teeth reddish brown; in lighter form, abdominal ventrites 2–5 darkened only in posterior half, pygidium only darkened at narrow anterior line, pro- and mesotibia yellowish brown; dull, with weak iridescent sheen; elytra lighter than pronotum, with a yellowish brown, narrow to broad zigzag band extending from primary costa 5 to 1, rarely from interstice V, sometimes very pale or interrupted; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.0; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface dense, large and coarse punctures. Frontoclypeal suture moderately incurved, distinct.An inverted triangular area on anterior half of frons punctate as clypeus, other portions with very dense and large punctures, smaller posteriad. Interocular distance/maximum head width approximately 0.65. Length of antennal club 0.9 times the combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third, slight concave in posterior third, moderately excurved at middle.Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line absent, other marginal lines complete. Lateral fovea indistinct. Disc with dense, rather large and transverse punctures, smaller and rounded along lateral and posterior margins. Lateral margin with several long setae.

Scutellum triangular, lateral margin moderately excurved, or semielliptical; with dense and small punctures, as well as a few erect and short setae.

Elytra. Surface weakly wrinkled, intervals moderately elevated, subsutural interstice widest; strial punctures dense and large, annulated, subsutural interstice with irregularly doubled secondary stria, interstices II–IV each with a somewhat irregular secondary stria, all intervals with moderately dense punctures, which are slightly or distinctly smaller than strial ones, each bearing a minute seta. Humeral and apical protuberance moderately prominent. Lateral carina slightly expanded from base to apical third. Epipleuron with a row of moderately dense and short setae from base to apical two fifths, usually partly abraded.

Abdomen. Propygidium with dense, transverse and fine striation. Pygidium with moderately dense and transverse striation, sparse only on the greatest tumidity; with sparse and long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites not carinate laterally; ventrites 2–5 with transverse and incompletely annulated punctures, short and moderately dense medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 densely rugostriolate; ventrites 2–5 each with a transverse and moderately dense row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron with sparse and small punctures, moderately dense and long setae. Ventral mesothoracic surface with dense and small punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and long setae, medial area with moderately dense and small punctures, setae slightly shorter. Metacoxa punctate as lateral metasternum but bearing sparser setae. Width of mesofemur/narrowest extremity of mesosternum approximately 8.0.

Legs. Protibia bidentate, both teeth distinct; apical tooth slender and extending to anterior margin of protarsomere 2, preapical tooth acute. Inner spur inserted at the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches equally as long as and two times wider than their upper branches; lower margin of lower branch of inner protarsal claw not convex next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 smaller. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of moderately dense and long setae, one lateral and the other submedial, also with some irregular and long setae sublaterally; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See fig. 23E–H.

Female. Usually entirely black, a vague lateral area of pronotum orange brown, each abdominal ventrite with a yellow spot at lateralmost area, antenna yellowish brown; elytra not paler than pronotum, zigzag band usually narrower or breaking up into three small spots. Rarely generally with similar color pattern like male. Antenna shorter, length of antennal club approximately 0.8 times the combined length of antennomeres 1–6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to anterior margin of protarsomere 2; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of protarsomere 5 distinctly smaller than in male; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch.

Measurement. Body length: 14.6–15.3 mm in males and 14.3–16.0 mm in females, greatest width: 7.8–8.9 mm in males and 7.5–9.4 mm in females.

Differential diagnosis. Anomala fenestrata sp. nov. is very similar to A. flavofasciata but larger. The parameres are stouter and have larger membranous areas laterally and ventrally. The ventral surface of the parameres is becoming abruptly membranous in the apical half.

Etymology. The name is derived from the Latin adjective fenestratus (having a window), alluding to the broad, translucent membranous structure present in the sclerotized parameres.