Anomala lhoba Zhao, Fujioka & Zorn, 2025

Zhao, Ming-Zhi, Fujioka, Masayuki & Zorn, Carsten, 2025, Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae), Zootaxa 5693 (3), pp. 301-356 : 316-318

publication ID

https://doi.org/10.11646/zootaxa.5693.3.1

publication LSID

lsid:zoobank.org:pub:3F45C3AC-27B2-4423-A260-E6586E0E27BA

DOI

https://doi.org/10.5281/zenodo.17322495

persistent identifier

https://treatment.plazi.org/id/4A5987BA-FFA9-930C-E9CA-84D4FEE1F846

treatment provided by

Plazi

scientific name

Anomala lhoba Zhao, Fujioka & Zorn
status

sp. nov.

Anomala lhoba Zhao, Fujioka & Zorn View in CoL , sp. nov.

[Chinese vernacular name: ḆƋ异丽aek]

( Figs 5D–F View FIGURE 5 , 11A View FIGURE 11 , 19E–H View FIGURE 19 )

Type material ( 19♂♂, 9♀♀). Holotype: ♂ ( SCAU) “ TIBET, Nyingchi, Medog, Bari , Renqinbeng , 1942m, 2014- VII-29 small lamp trap, Xiao-Dong Yang Leg. 14Y0324 CCCC” . Paratypes: 1♀ ( CCPC) “ TIBET, Nyingchi, Medog, Bari , Renqinbeng , 1942m, 2014-VIII-7 small lamp trap, Xiao-Dong Yang Leg. 14Y0415 CCCC” ; 1♂ ( CCPC) “ TIBET, Nyingchi, Medog, Bari , Renqinbeng , 1942m, 2014-VIII-7 small lamp trap, Xiao-Dong Yang Leg. 14Y0416 CCCC” ; 1♂ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Renqinbeng , 2023.VII.22, Zhi-Teng Chen leg.” ; 1♂ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Medog , 1117 m, 29.33116683°N, 95.33869959°E 2025.VI.28, Ke-Dian Chen leg.” GoogleMaps ; 1♂ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Bolonggong (80k), 2243 m, 29.65540554°N, 95.49091649°E 2023.VII.10, Hao-Yi Liu leg. // Voucher ACC1” GoogleMaps ; 3♂♂, 1♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill. , 1753 m, 29.214870°N, 95.169272°E, 18–30.VII.2023 (at light), Yao-Nan Zhang leg.” GoogleMaps ; 3♂♂, 1♀ ( ZYPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill. , 1753 m, 29.214870°N, 95.169272°E, 18–30.VII.2023 (at light), Yao-Nan Zhang leg.” GoogleMaps ; 3♀♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng, Gelin Vill. , 1768 m, 29.21315309°N, 95.17111894°E, 2023.VII.13-14, Hao-Yi Liu leg.” GoogleMaps ; 5♂♂ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge , S of Bolonggong, 2023.VII.9, Xu-Hong-Yi Zheng” ; 2♂♂, 2♀♀ ( ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge , 29.62527885°N, 95.48554466°E, 1681 m, 2023.VII.9, Zhi-Teng Chen leg.” GoogleMaps ; 1♀ ( SCAU) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge , 29.62527885°N, 95.48554466°E, 1681 m, 2023.VII.9, Zhi-Teng Chen leg. // ACC2” GoogleMaps ; 1♂ ( ZMPC) “ CHINA: TIBET, Nyingchi City , S of Medog Co., 1894 m 29.32160167N. 95.36277584E 2023.VII.10 Zhi-Teng Chen lgt.” GoogleMaps

Description. Male. Body shape elongated ovoid, weakly convex.

Color . Generally dark reddish brown to black; shiny, with strong bronze or greenish luster; elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1; antennae yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown.

Head. Greatest width/length of clypeus approximately 2.3; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface densely and coarsely rugopunctate. Frontoclypeal suture straight, distinct. An inverted triangular area on anterior third of frons punctate as clypeus, other portions with dense and large punctures, slightly sparser and smaller posteriad, sparse on vertex. Interocular distance/maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1–6. Inner margin of eye with a few moderately long setae.

Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third or posterior third, curved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae.

Scutellum triangular, lateral margin weakly to moderately excurved; with dense and large punctures.

Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive,irregular and large punctures,interstice II with a secondary stria, all intervals with moderately dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protberance strongly prominent. Lateral carina expanded from base to apical two fifths. Epipleuron with a row of dense and moderately long setae from base to near apex, often partly abraded.

Abdomen. Propygidium with very dense, transverse and fine striation. Pygidium with dense and transverse rugostriation, sparse around the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1–3 strongly carinate laterally; ventrites 2–5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2–5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin.

Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur/narrowest extremity of mesosternum approximately 6.0.

Legs. Protibia bidentate, both teeth distinct; apical tooth extending to anterior margin of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equal in mesotarsal claw; lower margin of lower branch of inner protarsal claw convex distally next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Protarsomeres distinctly thickened, each tarsomere 5 with an internomedial denticle, that of protarsomere 5 largest and triangular, others small and acute, metatarsomere 5 with an additional rounded basal protuberance. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae.

Male genitalia. See fig. 19E–H.

Female. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1–6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small and acute, basal protuberance of metatarsomere 5 weak; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw as long as its upper one and the lower margin not convex.

Measurement. Body length: 11.9–14.0 mm in males and 12.2–14.0 mm in females, greatest width: 6.4–8.0 mm in males and 7.1–7.9 mm in females.

Differential diagnosis. Anomala lhoba sp. nov. can be distinguished from all other species of the A. flavonotata - subgroup by the shortest upper branch of left paramere, as well as the simple lower branches of both parameres, which are not bifurcate in lateral view.

Etymology. The new species is named after the Lhoba people, an ethnic group traditionally residing in the Motuo region.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Anomala

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