Arge sogdiana Zhelochovtsev, 1976

27. Arge sogdiana Zhelochovtsev, 1976

( Figs 38 View FIGURE 38 , 39 View FIGURE 39 )

Arge sogdiana Zhelochovtsev, 1976: 59–60 , ♂, ♀); Taeger et al. 2010: 139.

Current status. Arge sogdiana Zhelochovtsev, 1976 .

Type locality. Tajikistan, Pamir Mountains, Khabost.

Holotype: ♀, <golden circle> // окр[естности]. Хорога, Хабост, [h=] 2100 [ Tajikistan, Pamir Mountains , vicinity of Khorog, Khabost, 37°46'N 71°60'E], 1.06.1956, Желоховцев [leg. A. V. Zhelochovtsev] // GoogleMaps Holotype Arge sogdiana sp. nov., ♀, A. Zhelochovtsev det. 1973 [ ZMMU] .

Paratypes: ♂, <golden circle>, окр[естности]. Хорога, Шугн. [Шуганский] хр[ебет]., [h=] 2500 [ Tajikistan, Pamir Mountains , vicinity of Khorog, Shuganskiy Ridge], 24.05.1956, Желоховцев [leg. A. V. Zhelochovtsev] // Allotype Arge sogdiana sp. nov., ♂, A. Zhelochovtsev det. 1973 [ ZMMU]; ♂, Гиссар[ский] хр[ебет]., Такоб [ское] уЩ[елье]. [ Tajikistan, Pamir Mountains, Hisar Range, Takob Gorge], 14.06.1952, Н. Филиппов [leg. N. Filippov] // Paratype Arge sogdiana sp. nov., ♂, A. Zhelochovtsev det. 1973 [ ZMMU]; ♀, Гиссарский хр[ебет]., Такобское уЩ[елье]. [ Tajikistan, Pamir Mountains , Hisar Range, Takob Gorge], 7.06.1952, Н. Филиппов [leg. N. Filippov] // Paratype Arge sogdiana sp. nov., ♀, A. Zhelochovtsev det. 1973 [ ZMMU]; ♀, Гиссарский хр[ебет]., АнЗобск[ий] пер[евал]. [h=] 2100 [ Tajikistan, Pamir Mountains, Hisar Range, Anzob Pass], 23.07.1953, А. Цветаев [leg. A. Tsvetaev] // Paratyp [e] Arge sogdiana sp. nov., ♀, A. Zhelochovtsev det. 1973 [ ZMMU]; ♀, Туркест[анский] хр[ебет]., с[еверный]. склон, уЩ[елье]. Кусавли [h=] 2000 [ Tajikistan, Pamir Mountains, Turkestan Ridge, northern slope, Kusavli Gorge], 7.07.1959, И. МалЯвин [leg. I. Malyavin] // Paratyp [e] Arge sogdiana sp. nov., ♀, A. Zhelochovtsev det. 1973 [ ZMMU]; ♀, Таджикистан, Памир, г[ород]. Хорог [ Tajikistan, Pamir Mountains, Khorog], 29.05.1968, В . Лоскот [leg. V. Loskot] // ботсад, выс. 2300 м, на цв. Ферулы Григорьева. [Botanical garden, h= 2300 m, on flowers of Ferula grigoriewii B. Fedtsch.]// Paratyp [e] Arge sogdiana sp. nov., ♀, A. Zhelochovtsev det. 1973 [ ZMMU] .

Description. Female ( Fig. 38 View FIGURE 38 ). Body length 8.7 mm; fore wing length 7.7 mm.

Colour. Head, thorax, two basal antennomeres, tergite 1 in part, except outer margin, legs black with slight blue metallic lustre; flagellum dark brown; abdomen yellowish-orange; tibiae and tarsi on fore and middle legs yellow-white (apical tarsomeres entirely darkened), hind tibia in basal half whitish. Mandibles in middle reddish, mouthparts brown. Wings hyaline, cell C yellowish, greyish towards apex. Apex of pterostigma, veins C and Sc, bases of other veins yellow, rest and pterostigma brown. Below pterostigma small indistinct dark spot or not. Pubescence yellowish-white.

Head. Head behind eyes slightly expanded. Distance between eyes 1.3 × vertical diameter of eye; eye with vertical diameter 1.3 × its horizontal diameter. Postocellar area slightly convex, with weak lateral furrows. POL: OCL: OOL as 1.4: 1.0: 1.8. Medial fovea broad, flat in lower part, with weak rounded pit in upper part, open towards frontal area. Lateral carinae sharp, converging downwards, not connected to each other. Supraclypeal area strongly roundly convex, evenly punctuated. Malar space smaller than diameter of frontal ocellus. Clypeus shiny, sparsely punctate, with medial semi-circular notch on ventral margin. Labrum with slightly concave ventral margin.

Antennae. Antennal length 1.4 × maximum head width; flagellum weakly curved at base, apex narrowly rounded; its setae short, dense, adhering.

Thorax. Mesopleurae in lower part with glabrous band; scutellum weakly convex, narrowly rounded at apex.

Legs. Middle tibia with preapical spurs; tibial spurs simple. Preapical spurs on hind tibiae absent. Basal tarsomeres of hind tarsus longer than next three combined.

Wings. In fore wing, cell 3Rs with anterior margin slightly longer than posterior margin, expanded to external margin. Vein 3r-m roundly curved. Vein B of fore wing joins vein R well before (longer than vein Sc) junction point of vein M. Membrane of cell 1Rs with very rare short setae. Wing margin between veins M and Cu non-ciliate, without glabrous band along margin.

Abdomen. Abdomen oval (in dorsal view), basal tergites glabrous. Saw sheath length 1.8 × its width in ventral view, simple, parallel, broadly rounded at apex (ventral view), on inner surface without blunt spines; in lateral view triangular, tapered at apex, apex rounded, slightly convex on ventral margin.

Lance in lateral view convex, medially with pronounced annuli. Maximum width of lance slightly greater than lancet width. Carina of apical part of dorsal margin weakly expressed, but with distinct irregular denticles, remaining part of dorsal margin with longitudinal deep fossa. Lancet with straight dorsal and convex medially ventral margin. Annuli nearly straight and dorsally inclined towards apex, in apical part indistinct, basally weakly curved. Annular spines (ctenidia) short, simple, several times shorter than width of annulus. Ventral margin with 23 well-distinct, strongly sclerotised serrulae. Serrulae triangular, oblique, with small irregular denticles, indistinct in apical part. Between serrulae two (less often three) unequal, straight pore canals of marginal sensilla.

Variability. Paratypes of females are variable: body length 8.7–8.9 mm; fore wing length 7.7–7.9 mm; whole tergite 1 black; legs black with strong or slight blue metallic lustre; only apexes of tarsomeres on fore and middle legs darkened; abdomen on tergites 2–6 with small, indistinct medial black spot; preapical spurs on hind tibiae developed, but 0.3 × as long as apical spurs or strongly reduced, equal in length to pubescence of tibiae and differing only in thickness and darker colour; lateral carinae converging downwards or parallel.

Male ( Fig. 39 View FIGURE 39 ). Body length 7.6 mm; fore wing length 7.5 mm.

In colouration as female, but body without distinct metallic lustre, on 5–6 basal tergites black medial spots decreasing to apex of abdomen. Legs completely black. Wings infuscated, to apex slightly lighter. Veins and pterostigma black. Pubescence of head, thorax and legs black. Distance between eyes 1.3 × vertical diameter of eye; eye with vertical diameter 1.6 × its horizontal diameter. Antennal length 1.7 × maximum head width; POL: OCL: OOL as 1.1: 1.0: 1.3. Flagellum weakly curved, pointed at apex, with long erect setae on front surface. Basal tarsomeres of hind tarsus shorter than next three combined. Scutellum broadly rounded. Subgenital plate in ventral view flat, slightly rounded on posterior margin. In fore wing, cell 3Rs with posterior margin slightly longer than anterior margin, expanded to external margin. Vein 3r-m roundly curved. Veins B and M of fore wing join up at same point on vein R. Membrane of cell 1Rs with dense short setae. Wing margin between veins M and Cu ciliate, length of setae equal to width of vein M.

Genitalia as in Fig. 39j View FIGURE 39 , gonostipes narrow, parallel. Harpe elongate, rounded at apex, 1.5 × as long as width at base, evenly covered by setae. Valve of penis subtriangular, with rounded lobe at apex. Valviceps in dorsal view slightly widened at apex, with small triangular lateral lobe, in middle with rounded lateral lobe; in lateral view apex in form of triangular, strongly sclerotised lobe, curved dorsally with large pores along external margin; ventrally with large, elongated, at apex rounded and weakly pigmented, poreless, curved at base lobe and notch at base. Dorsal process (ergot) short and broad at base.

Remark. The study of the type series made it possible to establish that the males ( paratypes) of A. sogdiana are identical to those of A. gussakovskiji . In the Arge ochropus species group, the males have a specific shape of valviceps, and females have sickle-shaped lancet with expanded annular spines. Females from this type series do not belong to the A. ochropus group, and strongly differ in colouration from males, as noted by Zhelochovtsev (1976) himself. The type locality of the males of A. sogdiana and A. gussakovskii coincide. The holotype of A. sogdiana is the female, so the status of this species remains valid and was not changed. The males are unknown.