Bactrocera ( Bactrocera ) wallacei Drew & Hancock, 2025

Bactrocera ( Bactrocera) wallacei Drew & Hancock , new species

( Fig. 1 View Fig )

Bactrocera ( Bactrocera) epicharis View in CoL : Drew & Romig, 2013: 79–80; 2016: 151; not Hardy, 1970: 119. Misidentification.

Type material. Holotype: male, INDONESIA: North Moluccas [ North Maluku] – Maluku, North Maluku, West Halmahera, Goal , 29.v.2007, coll. Raais Abdullah, attracted to cue lure in forest, deposited in Queensland Museum ( QMIC), Brisbane . Paratypes: 2 males, 5.vi.2007, same label data as holotype ; 1 male ( 3.vi.2007), 1 male ( 10.vi.2007), North Maluku, Kodya Ternate, Foramadiahi, coll. La Ruti, attracted to cue lure in forest . Paratypes in Department of Agriculture and Fisheries Collection ( QDPC), Brisbane .

Diagnosis. A large species (wing 7.5 mm); face fulvous with a pair of medium-sized to large circular black spots; postpronotal lobes and notopleural calli yellow; scutum black; lateral postsutural yellow vittae present, elongate and parallel-sided; medial postsutural yellow vitta absent; no yellow spot anterior to notopleural suture; anepisternal (mesopleural) stripe reaching midway between anterior margin of notopleuron and anterior notopleural seta dorsally; scutullum yellow; wing with cells bc and c fuscous, microtrichia in anteroapical corner of cell c only; a broad dark fuscous costal band becoming paler between R 2+3 and R 4+5 and remaining of uniform width throughout; a broad dark fuscous anal streak; abdominal terga III–V entirely black.

Description. Male. Head: Height 2.0 mm. Frons length 1.57 times breadth, fuscous with fulvous along lateral and ventral margins and dark fuscous on anteromedial hump; orbital setae black: 1 s.or., 2 i.or.; lunule dark fuscous. Ocellar triangle black. Vertex fuscous. Face fulvous with a pair of medium-sized to large circular black spots; length 0.58 mm. Genae red-brown, dark fuscous subocular spot present; black seta present. Occiput fuscous, red-brown along eye margins; occipital row with 5–8 strong black setae. Antennae with segments 1 and 2 red-brown, segment 3 red-brown with fuscous on apex and outer surface; length of segments: 0.24 mm; 0.44 mm; 1.06 mm.

Thorax: Scutum black with dark fuscous below and behind lateral postsutural vittae, around notopleural suture, along lateral margins between postpronotal lobe and notopleuron, inside postpronotal lobe. Pleural areas entirely black. Yellow markings as follows: postpronotal lobes; notopleural calli; anepisternal stripe reaching midway between anterior margin of notopleuron and anterior notopleural seta dorsally, continuing to katepisternum as a small transverse spot, anterior margin slightly convex; anatergite (posterior apex black); anterior ⅔ katatergite (remainder black); two broad parallel-sided lateral postsutural vittae ending just behind ia. seta. Postnotum black. Scutellum yellow except for narrow black basal band. Setae (pairs): sc. 1; prsc. 1; ia. 1; p.sa. 1; a.sa. 1; mpl. 1; npl. 2; scp. 2.

Legs: All femora entirely fulvous; fore and hind tibiae dark fuscous, mid tibiae fuscous; fore tarsi with basal segment fulvous and apical four segments red-brown, mid and hind tarsi with all segments entirely fulvous; mid tibiae each with an apical black spur.

Wings: Length 7.5 mm; cells bc and c fuscous, microtrichia in anteroapical corner of cell c only; remainder of wings colourless except dark fuscous cell sc, dark fuscous costal band confluent with R 4+5 but tending fuscous between R 2+3 and R 4+5 and remaining of uniform width throughout, a broad dark fuscous anal streak; a dense aggregation of microtrichia around A 1 +CuA 2; supernumerary lobe of medium development.

Abdomen: Oval; terga free; pecten present on tergum III. Tergum I and sterna I and II wider than long. Tergum I black with a narrow transverse red-brown band across posterior margin but not reaching lateral margins; tergum II black with a narrow transverse fulvous band across posterior margin but not reaching lateral margins; terga III–V entirely black. A pair of oval black shining spots on tergum V. All sterna dark fuscous to black. Posterior lobe of surstylus short, sternum V with a deep concavity on posterior margin.

Female. No known record.

Male attractant. Cue lure.

Distribution. Known only from Halmahera and Ternate, North Maluku, Indonesia.

Host plants. No known record.

Etymology. This species is named after Alfred Russel Wallace in recognition of his pioneering field work in Wallacea and beyond.

Remarks. Bactrocera ( Bactrocera) wallacei was misidentified as B. epicharis ( Hardy, 1970) by Drew & Romig (2013, 2016), who also overlooked the description and discussion of B. epicharis based on new material from the Solomon Islands ( Drew & Romig, 2001). It is included here in the trivialis group of the nigella complex and most resembles B. atrabifasciata Drew & Romig, 2001 from the Solomon Islands, differing from it in the narrower anepisternal stripe and fuscous tibiae, and from both B. illusioscutellaris Drew & Romig, 2013 from Bhutan and Vietnam and B. trivialis ( Drew, 1971) from New Guinea in the broad costal band and darker abdomen.