PSAMMOBIIDAE J. Fleming, 1828

Family PSAMMOBIIDAE J. Fleming, 1828 View in CoL

Capsella – Deshayes, 1855c: 347-350, non J. E. Gray, 1851. — Type species ( SD herein): Capsa ( Capsella) crassula Deshayes, 1855 . — Current status: this is the same type species as that of Crassulobia Willan, 1993, so Deshayes genus falls into its synonymy. It is possible that Deshayes merely misused Gray’s genus, but that seems unlikely in that Gray’s genus belongs in the Donacidae and is now regarded as a synonym of Donax Linnaeus, 1758 .

REMARK

Capsa Bruguière, 1797 View in CoL , was misused for some of the following species, but it is a tellinid genus. Recently, the long-used genus Soletellina Blainville, 1824 , was synonymized with Hiatula Modeer, 1793 View in CoL ( Matsubara, 2013).

abrupta, Psammobia – Deshayes, 1855c: 324-325. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984300, holotype. — Current status: synonym of Gari pulcherrima (Deshayes, 1855) View in CoL ( Willan 1993: 35, fig. 141 – holotype).

amoena , Psammobia – Deshayes, 1855c: 323. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1964010, holotype. — Current status: synonym of the Asian Gari amethystus (W. Wood, 1815) [ Solen ] ( Willan 1993: 38- 39, fig. 152 – holotype).

angusta View in CoL , Psammobia – Deshayes, 1855c: 320. — Type locality: Senegal; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984288.1, lectotype ( Willan 1993); 1984288/2, paralectotype. — Current status: synonym of Gari modesta (Deshayes, 1855) View in CoL [ Psammobia ] (E. A. Smith 1885: 95; Willan 1993: 23, fig. 84 – lectotype).

anomala View in CoL , Psammobia – Deshayes, 1855c: 320. — Type locality: Zebu, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984290.1 lectotype ( Willan 1993); 1984290/2- 5 paralectotypes. — Current status: Gari anomala (Deshayes, 1855) View in CoL ( Willan 1993: 22-23, figs 7, 8, 76 [ lectotype]-81; Huber 2010a: 324-325; Willan & Tagaro, in Poppe 2011: 356, pl. 1173, figs 1-2; Matsukuma, in Okutani 2017: 1260, pl. 554, fig. 5).

basteroti View in CoL , Tellina View in CoL – Deshayes, 1857-DescrCoqFoss: 348. Based on Tellina biangularis Deshayes View in CoL var. “β” of Basterot (1825: 86). — Type locality: Bordeaux. Miocene (Aquitanian). Synonym of Solecurtus affinis ( Dujardin, 1837) View in CoL (d’Orbigny 1852: 99, no. 1853). Gari affinis ( Dujardin, 1837) ( Glibert & Van de Poel 1967: 122) View in CoL . — Current status: Gari ( Psammobia) affinis ( Dujardin, 1837) View in CoL ( Lesport et al. 2019: 44, pl. 17, figs 6-9).

baudoni , Psammobia –Deshayes, 1857-DescrCoqFoss: 378-379, 23, pl. 25, figs 15-18 [in text as pl. 25]. — Type locality: Hadancourtle-Haut-Clocher (Oise). — Type age: Eocene (Lutetian). — Current status: Gari ( Psammotaena) baudoni (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93). bicarinata View in CoL , Psammobia – Deshayes, 1855c: 322. — Type localities: Madagascar & Zanzibar; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984297, 2 syntypes. — Current status: Gari bicarinata (Deshayes, 1855) View in CoL ( Oliver 1992: 163, pl. 36), or synonym of Psammobia insignis Deshayes, 1855 ( Willan 1993: 37; Rusmore-Villaume 2008: 260-261).

caillati View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 381, 23, pl. 23, figs 19-20. — Type localities: Parnes, Thiverval-Grignon, Chaumont-en-Vexin & Gomerfontaine. — Type age: Eocene (Lutetian). — Type species (OD) of Psammodonax Cossmann, 1886 View in CoL . Psammodonax caillati (Deshayes, 1857) ( Glibert & Van de Poel 1967: 120) View in CoL . — Current status: Gari ( Psammodonax) caillati (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

callosa View in CoL , Sportella View in CoL – Deshayes, 1857-DescrCoqFoss: 48, pl. 48, figs 18-20; 1858-DescrCoqFoss: 595-596. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). Gobraeus callosus (Deshayes, 1857) ( Glibert & Van de Poel 1967: 124) . — Current status: Gari ( Psammotaena) callosa (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

candidula View in CoL , Psammobia – Deshayes, 1855c: 319. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984286, 3 syntypes. — Current status: Gari candidula (Deshayes, 1855) View in CoL ( Huber 2015: chapter 5 on CD).

chinensis View in CoL , Capsa ( Capsella) – Deshayes, 1855c: 348. — Type locality: China; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984261, 2 syntypes. — Current status: Gari ( Psammotaena) chinensis (Deshayes, 1855) View in CoL ( Willan 1993: 60; Huber 2010a: 329, 695).

compta , Psammobia – Deshayes, 1855c: 321. — Type locality: Tasmania; Cuming coll. — Type age: Recent. — Type material: NHMUK 1841.2.6.423, lectotype ( Willan 1993); 1837.7.15.310, paralectotype; 1837.7.15.144, paralectotype. — Current status: synonym of Gari livida ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 41-42, fig. 162 – lectotype).

consobrina View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 373-374, 23, pl. 23, figs 3-4. — Type localities: Brimont & Châlons-sur-Vesle. — Type age: Paleocene (Thanetian). 1858: 552. Garum consobrina (Deshayes, 1857) ( Glibert & Van de Poel 1967: 119) . — Current status: Gari ( Garum) consobrina (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Leroy et al. 2014: 28, pl. 12, fig. 1a-b).

contraria View in CoL , Psammobia – Deshayes, 1863: 11, pl. 1, figs 20-21. — Type locality: La Réunion. — Type age: Recent. — Type material: missing ( Willan 1993: 9). — Type species (SD Oliver, 1992) of Heteroglypta Martens, 1880 View in CoL . — Current status: Heteroglypta contraria (Deshayes, 1863) View in CoL ( Oliver 1992: 162, pl. 36, as Gari ( Heteroglypta) ; Willan 1993: 9-11, figs 27-37, 382; Huber, 2010a: 332; Willan & Tagaro, in Poppe 2011: 362, pl. 1176, figs 4-5; Matsukuma, in Okutani 2017: 1262, pl. 555, fig. 5; Lutaenko et al. 2021: 220).

corrugata View in CoL , Psammobia – Deshayes, 1855c: 324. — Type locality: Zebu Island, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 196413, holotype. — Current status: synonym of Gari maculosa ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 26-27, fig. 102 – holotype).

crassula View in CoL , Capsa ( Capsella) – Deshayes, 1855c: 349. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984268.1, lectotype ( Willan 1993); 1984268/2-3, paralectotypes. — Type species (OD) of Crassulobia Willan, 1993. — Current status: Gari ( Crassulobia) crassula (Deshayes, 1855) View in CoL ( Willan 1993: 68-70, figs 289 [ lectotype]-297, 404-405; Huber 2010a: 328). debilis View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 377-378, 22, as P. fragilis View in CoL , pl. 22, figs 15-17; 1865-DescrCoqFoss: 666 [error corrected]. — Type locality: Châlons-sur-Vesle. — Type age: Paleocene (Thanetian). 1858: 552. Gobraeus debilis (Deshayes, 1857) ( Glibert & Van de Poel 1967: 124) . — Current status: Gari ( Psammotaena) debilis (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Leroy et al. 2014: 28, pl. 12, figs 2-4).

difficilis View in CoL , Capsa ( Capsella) – Deshayes, 1855c: 347. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984262.1, lectotype ( Willan 1993); 1984268/2-3, paralectotypes; 1984262, 3 syntypes. — Current status: synonym of Gari ( Psammotaena) elongata ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 61-62, fig. 251 – lectotype).

dispar View in CoL , Psammobia – Deshayes, 1855c: 325. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1985170.1, lectotype ( Willan 1993); 1985170/2, paralectotype. Gari ( Heteroglypta) dispar (Deshayes, 1855) View in CoL ( Oliver 1992: 164, pl. 36). — Current status: synonym of Gari pennata (Deshayes, 1855) View in CoL ( Willan 1993: 31, fig. 120 – lectotype; Rusmore-Villaume 2008: 262-263).

donacilla View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 382, 23, as Psammobia donacina , pl. 23, figs 21-22; 1865-DescrCoqFoss: 666 [error in pl. expl. corrected]. — Type localities: Thiverval-Grignon, Parnes, Damery, Chaumont-en-Vexin, Fontenay-Saint-Père, Boursault & Gomerfontaine. — Type age: Eocene (Lutetian). — Current status: Gari ( Psammodonax) donacilla (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

dutemplei View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 374, 23, pl. 23, figs 10-12. — Type locality: Damery. — Type age: Eocene (Lutetian). — Type species (OD) of Garum Dall, 1900 . Garum placed on Official List by ICZN Opinion, 910 (1970), but listed there with the wrong type species [ Psammobia filosa Conrad, 1833 ]. Garum dutemplei (Deshayes, 1857) ( Glibert & Van de Poel 1967: 119) . Gari ( Psammodonax) dutemplei (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70) View in CoL , – Current status: Gari ( Garum) dutemplei (Deshayes, 1857) ( Pacaud 2008: 93) View in CoL .

elegans View in CoL , Psammobia – Deshayes, 1855c: 322. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984298, 6 syntypes. — Current status: synonym of the western Indo-Pacific Psammobia insignis Deshayes, 1855 ( Willan 1993: 37).

grata , Psammobia – Deshayes, 1855c: 318-319. — Type locality: “ Amboina ”; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984283, holotype. — Current status: synonym of the northeastern Atlantic Gari ( Gobraeus) depressa (Pennant, 1777) [ Tellina ] ( Willan 1993: 49).

insignis , Psammobia – Deshayes, 1855c: 322. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984296, holotype. Gari insignis (Deshayes, 1855) ( Willan 1993: 37) View in CoL . — Current status: the western Indo-Pacific Gari ( Psammobia) insignis (Deshayes, 1855) ( Huber 2010a: 326) View in CoL .

intermedia View in CoL , Psammobia – Deshayes, 1855c: 319. — Type locality: Portugal; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984284, 2 syntypes. Synonym of Gari ( Gobraeus) virgata ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 49). — Current status: Gari intermedia (Deshayes, 1855) View in CoL ( Cossignani & Ardovini 2011: 57, 471).

lamarckii , Psammobia – Deshayes, 1857-DescrCoqFoss: 376; first edition, pl. 2, figs 24, 25 [in text as 24, 26]; 1865-DescrCoqFoss: 665 [error noted]. — Type localities: Parnes, Mouchy-le-Châtel, Montmirail & Vaudancourt. — Type age: Eocene (Lutetian). — Type species (M) of Amphipsammus Cossmann, 1913. Gobraeus lamarckii (Deshayes, 1857) , as G. “ lamarcki View in CoL ”) ( Glibert & Van de Poel 1967: 125). — Current status: Gari ( Amphipsammus) lamarckii (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 94).

lata View in CoL , Psammobia – Deshayes, 1855c: 318. — Type localities: Central America & Santa Elena, Ecuador; Cuming coll. — Type age: Recent. — Type material: NHMUK 1966549, lectotype, 2 paralectotypes; 1984279, probable paralectotype. Gari ( Gobraeus) lata (Deshayes, 1855) ( Huber 2010a: 327) View in CoL . — Current status: Gari lata (Deshayes, 1855) ( Coan & Valentich-Scott 2012: 685) View in CoL .

layardi View in CoL , Capsa ( Capsella) – Deshayes, 1855c: 348. — Type locality: Ceylon; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984276, 3 syntypes. — Current status: Gari ( Psammotaena) layardi (Deshayes, 1855) ( Huber 2010a: 329) View in CoL .

layardi View in CoL , Psammobia –Deshayes, 1855c: 323.— Type locality: Philippine Islands;Cuming coll.— Type age:Recent. —Type material: NHMUK 196422, holotype. — Current status: synonym of Gari maculosa ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 26-27, fig. 99 – holotype).

lunulata View in CoL , Capsa ( Capsella) – Deshayes, 1855c: 349. — Type localities: Philippine Islands; Ceylon (var.); Cuming coll. — Type age: Recent. — Type material: NHMUK 1984272, 5 syntypes. — Current status: Soletellina lunulata (Deshayes, 1855) ( Willan 1993: 75; Huber 2010a: 330), now Hiatula lunulata (Deshayes, 1855) View in CoL .

marmorea, Psammobia – Deshayes, 1855c: 324. — Type localities: Darnley Island, Sydney, Australia & Moluccas (latter two “var.); Cuming coll. — Type age: Recent. — Type material: NHMUK 1846.9.16.112, holotype; 1846.9.14.111, syntype of variety Alpha from Sydney; 196447-196448, syntypes of variety Beta from Moluccas. — Current status: synonym of Gari maculosa ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 26-27, fig. 103 – holotype).

maxima View in CoL , Psammobia – Deshayes, 1855c: 317. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1966552.1-2, lectotype & paralectotype. — Current status: the Panamic Gari ( Gobraeus) maxima (Deshayes, 1855) View in CoL ( Huber 2010a: 326; Coan & Valentich-Scott 2012: 685).

menkeana, Psammobia – Deshayes, 1855c: 319-320. — Type locality: east coast of Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984287.1, lectotype ( Willan 1993); 1984287/2-3, paralectotypes. — Current status: synonym of Gari modesta (Deshayes, 1855) View in CoL [ Psammobia ] (E. A. Smith 1885: 95; Willan 1993: 23, fig. 83 – lectotype).

minima , Capsa View in CoL –Deshayes, 1857-DescrCoqFoss: 386, 11 bis, pl. 11 bis, figs 29-31 [in text as pl. 16 bis, figs 81-33]; 1865-DescrCoqFoss: 665 [errors noted]. — Type localities: Montjavoult, Thiverval-Grignon, Houdan & Coincourt. — Type age: Eocene (Lutetian). — Type species (OD) of Asaphinella Cossmann, 1886 View in CoL . — Current status: Asaphinella minima (Deshayes, 1857) View in CoL ( Glibert & Van de Poel 1967: 126; Le Renard & Pacaud 1995: 71; Pacaud 2008: 93; Nevesskaja et al. 2013: 377, fig. 138-5).

minor View in CoL , Capsa ( Capsella) – Deshayes, 1855c: 347. — Type locality: Manilla, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984269.1, lectotype ( Willan 1993); 1984269/2-4, 3 paralectotypes. Synonym of Gari ( Psammotaena) elongata ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 61-62, fig. 250 – lectotype). — Current status: Psammotaena minor (Deshayes, 1854) ( Qi 2004: 292, pl. 159, fig. A).

modesta View in CoL , Psammobia – Deshayes, 1855c: 319. — Type locality: Moreton Bay, Queensland, Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984285.1, lectotype ( Willan 1993); 1984285/2-3, paralectotypes. — Current status: Gari modesta (Deshayes, 1855) View in CoL ( Willan 1993: 23-26, figs 82 [ lectotype]-93, 383). neglecta View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 375-376, 23, pl. 23, figs 5-7. — Type locality: Ludes. — Type age: Eocene (Priabonian). Gobraeus neglectus (Deshayes, 1857) ( Glibert & Van de Poel 1967: 125) . — Current status: Gari ( Psammotaena) neglecta (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Pacaud & Ledon 2007: 11).

nitens View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 380, 21, pl. 21, figs 27-28. — Type locality: Ormoy-la-Rivière. — Type age: Oligocene (Rupelian). — Current status: Gari nitens (Deshayes, 1857) ( Lozouet & Maestrati 2012a: 258, 261, fig. 166: 5-8).

nitida View in CoL , Psammobia –Deshayes, 1857-DescrCoqFoss: 370, 24, pl. 24, figs 1-2. — Type localities: Parnes, Thiverval-Grignon, Houdan, Hermonville & Coincourt. — Type age: Eocene (Lutetian). Synonym of Soletellina appendiculata ( Lamarck, 1806) [ Solen View in CoL ] ( Cossmann 1886a: 89-90). — Current status: Gari ( Psammoica) appendiculata ( Lamarck, 1806) View in CoL ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

nivosa, Psammobia – Deshayes, 1855c: 317-318. — Type locality: Negros, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984277.1, lectotype ( Willan 1993); 1984277/2, paralectotype. — Current status: synonym of Gari ( Dysmea) occidens (Gmelin, 1791) View in CoL [ Solen View in CoL ] ( Willan 1993: 56-57, fig. 228 – lectotype).

oblonga , Psammobia – Deshayes, 1855c: 321. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984295, holotype. — Current status: synonym of the Red Sea Soletellina rosea (Gmelin, 1791) [ Solen ] ( Huber 2010a: 331, 695), now Hiatula rosea (Gmelin, 1791) .

obtusalis View in CoL , Donax View in CoL – Deshayes, 1825-DescrCoqFoss: 109-110, 9, pl. 18, figs 7-8; 1830b-EncyMeth: 101; 1835-HistNatAnim2: 251- 252; 1839-HistNatAnim3: 583; 1857-DescrCoqFoss: 383-384, as Psammobia obtusalis . — Type localities: Pierrelaye & Beauchamp. — Type age: Eocene (Bartonian). Psammodonax obtusalis (Deshayes, 1825) View in CoL ( Glibert & Van de Poel 1967 120). — Current status: Gari ( Psammodonax) obtusalis (Deshayes, 1825) ( Le Renard & Pacaud 1995: 70) View in CoL .

oriens View in CoL , Psammobia – Deshayes, 1855c: 318. — Type locality: Japan; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984277.1, lectotype ( Willan 1993); 1984277/2, paralectotype. Gari ( Gobraeus) oriens (Deshayes, 1855) View in CoL ( Willan 1993: 52-55, figs 214-216 [ lectotype]-222, 395; Willan & Tagaro, in Poppe 2011: 350, pl. 1170, figs 4-5, who treated it as a subspecies of the West African Gari castrensis Spengler, 1794 View in CoL ). Gari ( Gobraeus) oriens (Deshayes, 1855) ( Huber 2010a: 327) View in CoL . — Current status: Gobraeus oriens (Deshayes, 1855) (Matsukuma, in Okutani 2017: 1261-1262, pl. 555, fig. 2). Poppe (2018: 173) concluded that Gari oriens (Deshayes, 1855) View in CoL should be used for the Indo-Pacific species, with G. castrensis (Spengler, 1794) View in CoL restricted to the tropical eastern Atlantic.

ornata View in CoL , Psammobia – Deshayes, 1855c: 323. — Type locality:Ticao Island, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 196423, lectotype ( Willan 1993); 196424/1- 2, paralectotypes; 196425, 1 paralectotype; 196426, 2 paralectotypes. — Current status: synonym of Gari maculosa ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 26-27, fig. 100 – lectotype).

pallida View in CoL , Psammobia – Deshayes, 1855c: 323-324. — Type locality: Red Sea; Cuming coll. — Type age: Recent. — Type material: NHMUK 1964046.1, lectotype ( Willan 1993); 1964046/2, paralectotype; 1844.6.3.594, paralectotype. — Current status: Gari pallida (Deshayes, 1855) View in CoL ( Willan 1993: 19-22, figs 6, 7, 57 [ lectotype]-75; Huber 2010a: 325; Willan & Tagaro, in Poppe 2011: 356, pl. 1173, figs 3-4; Matsukuma, in Okutani 2017: 1260-1261, pl. 554, fig. 6).

palmula , Psammobia – Deshayes, 1855c: 325-326. — Type locality: Sydney, Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1985171, holotype. — Current status: synonym of Gari squamosa ( Lamarck, 1818) View in CoL [ Psammobia ] ( Hedley 1913: 274; Willan 1993: 33, fig. 133 – lectotype), or separable Grammatomya palmula (Deshayes, 1855) (Matsukuma, in Okutani 2017: 1261, pl. 554, fig. 9, as G. “ palmura ”).

papyracea View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 372. Replacement name for Psammobia tellinella (Deshayes, 1824) [ Solen View in CoL ] (see below), non P. tellinella Lamarck, 1818 .

pennata View in CoL , Psammobia – Deshayes, 1855c: 325. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material:NHMUK 1985169, holotype. — Current status: the Indo-Pacific Gari pennata (Deshayes, 1855) View in CoL ( Willan 1993: 31-33, figs 119 [ holotype]-130, 386; Huber 2010a: 325; Severns 2011: 480, pl. 220, fig.3; Willan &Tagaro, in Poppe 2011: 362, pl. 1176, figs 1-3; Boutet et al. 2020: 661).

petalina , Psammobia – Deshayes, 1855c: 320-321. — Type locality: China; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984292.1, lectotype ( Willan 1993); 1984292/2-3, paralectotypes. — Current status: Soletellina petalina (Deshayes, 1855) ( Willan 1993: 82-83, figs 331 [ lectotype]-335, 411; Huber 2010a: 331; Matsukuma, in Okutani 2017: 1262, pl. 555, fig. 8), now Hiatula petalina (Deshayes, 1855) View in CoL .

praestans View in CoL , Psammobia – Deshayes, 1855c: 322-323. — Type locality: Moluccas Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 196416, lectotype ( Willan 1993); 196417, paralectotype. — Current status: synonym of Gari maculosa ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 26-27, fig. 98 – lectotype).

puella, Psammobia – Deshayes, 1855c: 320. — Type locality: Australia; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984289, holotype.— Current status:synonym of Gari livida ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 41-42, fig. 161 – holotype).

pulcherrima View in CoL , Psammobia – Deshayes, 1855c: 325. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1985168, holotype. — Current status: the Indo-Pacific Gari pulcherrima (Deshayes, 1855) View in CoL ( Willan 1993: 35-37, figs 140 [ holotype]-148, 388; Huber 2010a: 325; Willan & Tagaro, in Poppe 2011: 358, pl. 1174, figs 5-8; Matsukuma, in Okutani 2017: 1261, pl. 554, fig. 10).

purpurea View in CoL , Sanguinolaria View in CoL – Deshayes, 1855c: 346. — Type locality: none given [clarified as Guaymas, Mexico, byCoan 2002: 4]; Cuming coll. — Type age: Recent. — Type material: NHMUK 1966539.1- 3, lectotype & 2 paralectotypes. — Current status: synonym of the Panamic Sanguinolaria tellinoides A. Adams, 1850 View in CoL ( Coan & Valentich-Scott 2012: 693).

radiata , Capsa ( Capsella) – Deshayes, 1855c: 348. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984267.1, lectotype ( Willan 1993); 1984267/2-3, paralectotypes. — Current status: synonym of Gari ( Psammotaena) elongata ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 61-62, fig. 253 – lectotype).

rosacea View in CoL , Capsa ( Capsella) – Deshayes, 1855c: 348. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984275.1 lectotype ( Willan 1993); 1984275/2-3, paralectotypes. — Current status: synonym of Gari ( Psammotaena) elongata ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 61-62, fig. 254 – lectotype).

rubicunda, Psammobia – Deshayes, 1855c: 324. — Type locality: Ticao Island , Philippine Islands ; Cuming coll. — Type age:Recent. — Type material: NHMUK 1984299.1, lectotype ( Willan 1993);

1984299/2, paralectotype. Synonym of Gari maculosa ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 26, fig. 101 – lectotype).

rufa View in CoL , Capsa ( Capsella) –Deshayes, 1855c: 347-348. — Type locality: Manilla, Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984263.1, lectotype ( Willan 1993); 1964263/2, paralectotype. — Current status: synonym of Gari ( Psammotaena) elongata ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 61-62, fig. 252 – lectotype).

solenella, Capsa ( Capsella) –Deshayes, 1855c: 349-350.— Type locality: Manilla, Philippine Islands; Cuming coll. — Type age: Recent. —Type material:NHMUK 1984271.1, lectotype ( Willan 1993); 1984271/2-3, paralectotypes.— Current status: synonym of Gari elongata ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 61-62, fig. 255 – lectotype).

spathula View in CoL , Psammobia –Deshayes, 1857-DescrCoqFoss: 382-383, 24, pl. 24, figs 3-5. — Type locality: Chaumont-en-Vexin. — Type age: Eocene (Lutetian). — Current status: Gari ( Psammodonax) spathula (Deshayes, 1857) View in CoL ( Le Renard & Pacaud 1995: 70; Pacaud 2008: 93).

stampinensis , Psammobia – Deshayes, 1857-DescrCoqFoss: 378, 23, pl. 23, figs 13-14. — Type localities: Morigny-Champigny, Etréchy. — Type age: Oligocene (Rupelian). Haesslet, Belgium; Délémont, Switzerland. — Current status: Psammobia stampinensis Deshayes, 1857 ( Lozouet & Maestrati 2012a: 258, 261, figs 166: 1-4).

striata View in CoL , Psammobia –Deshayes, 1855c: 321. — Type locality: Tasmania; Cuming coll. — Type age: Recent. — Type material: NHMUK 1842.11.2.8.9, 2 possible syntypes. — Current status: synonym of Gari livida ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 41-42, fig. 163 – possible syntype).

tellinella , Solen View in CoL – Deshayes, 1824-DescrCoqFoss: 28, 2, pl. 4, figs 1-2, non Psammobia tellinella Lamarck, 1818 ; 1832-EncyMeth: 963; 1857-DescrCoqFoss: 372, Psammobia papyracea , replacement name, when he moved this Solen View in CoL into Psammobia . — Type locality: Tancrou. — Type age: Eocene (Bartonian). Macropsammus tellinella (Deshayes, 1824) ( Glibert & Van de Poel 1967: 126) . Gari ( Psammoica) tellinella (Deshayes, 1824) ( Le Renard & Pacaud 1995: 70) View in CoL . — Current status: because Lamarck’s species is now placed in Gari View in CoL , this should once again be Gari ( Psammoica) papyracea (Deshayes, 1857) .

tenera View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 372, 24, pl. 24, figs 6-8. — Type localities: Damery, Parnes & Hermonville. — Type age: Eocene (Lutetian). Synonym of Soletellina appendiculata ( Lamarck, 1806) [ Solen View in CoL ] ( Cossmann 1886a: 89-90). — Current status: Gari ( Psammoica) appendiculata ( Lamarck, 1806) ( Le Renard & Pacaud 1995: 70) View in CoL .

tenuicula View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 380-381, 24, as P. “ tenuis View in CoL ”, pl. 24, figs 12-14; 1866: 666 [error corrected]. — Type localities: Gentilly, Hermonville & La Chapelle-en-Serval. — Type age: Eocene (Bartonian). Gobraeus tenuiculus (Deshayes, 1857) ( Glibert & Van de Poel 1967: 125) . — Current status: Gari ( Psammotaena) tenuicula (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70) View in CoL .

tenuis View in CoL , Capsa ( Capsella) – Deshayes, 1855c: 349. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984274.1, lectotype ( Willan 1993); 1984274/2, paralectotype. Soletellina tenuis (Deshayes, 1855) ( Willan 1993: 76) , now genus Hiatula View in CoL . — Current status: synonym of Gari virescens (Deshayes, 1855) View in CoL ( Huber 2015: chapter 5 on CD).

tenuis , Psammobia – Deshayes, 1855c: 320. — Type localities: Philippine Islands & China (var.); Cuming coll. — Type age: Recent. — Type material: NHMUK 1984291.1/2, lectotype & paralectotype. — Current status: synonym of Gari anomala (Deshayes, 1855) View in CoL ( Willan 1993: 22, fig. 77 – lectotype). togata , Psammobia – Deshayes, 1855c: 318. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984280.1, lectotype; 1984280/2, paralectotype. Gari ( Psammotaena) togata (Deshayes, 1855) View in CoL ( Willan 1993: 64-66, figs 269 [ lectotype]-274, 401). — Current status: Psammotaena togata (Deshayes, 1854) ( Qi 2004: 292, pl. 159, fig. D; Huber 2010a: 329; Willan & Tagaro, in Poppe 2011: 346, pl. 1168, fig. 6; Matsukuma, in Okutani 2017: 1261, pl. 554, fig. 15).

tripartita , Psammobia – Deshayes, 1855c: 321-322. — Type locality: Philippine Islands; Cuming coll. — Type age: Recent. — Type material: NHMUK 196414, lectotype ( Willan 1993); 196415/1-2, paralectotypes; 196416, paralectotype. — Current status: synonym of Gari amethystus (W. Wood, 1815) View in CoL [ Solen View in CoL ] ( Willan 1993: 38-39, fig. 151 – lectotype).

tristis View in CoL , Psammobia – Deshayes, 1855c: 318. — Type locality: Amboina ; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984282.1, lectotype ( Willan 1993); 1984272/2, paralectotype. — Current status: synonym of Gari ( Gobraus) strangeri (J. E. Gray, 1843) [ Psammobia ] ( Willan 1993: 49-50, fig. 206 – lectotype).

vaudini View in CoL , Psammobia – Deshayes, 1857-DescrCoqFoss: 383, 22, pl. 22, figs 12-14. — Type localities: Laon & Hérouval. — Type age: Eocene (Ypresian). Psammodonax vaudini (Deshayes, 1857) ( Glibert & Van de Poel 1967: 120) View in CoL . — Current status: Gari ( Psammodonax) vaudini (Deshayes, 1857) ( Le Renard & Pacaud 1995: 70) View in CoL .

virescens View in CoL , Capsa ( Capsella) –Deshayes, 1855c: 349. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1984270, 3 syntypes. Soletellina virescens (Deshayes, 1855) ( Willan 1993: 76; Qi 2004: 292-293, pl. 159, fig. B), now genus Hiatula View in CoL . — Current status: the southeast Asian Gari ( Psammotaena) virescens (Deshayes, 1855) ( Huber 2010a: 329) View in CoL .

vitrea , Sanguinolaria View in CoL – Deshayes, 1855c: 326. — Type locality: none given; Cuming coll. — Type age: Recent. — Type material: NHMUK 1985172, 2 syntypes. — Current status: Sanguinolaria vitrea Deshayes, 1855 View in CoL ( Huber 2010a: 332-333, 696; 2015: 749) from Madagascar .

zelandica View in CoL , Psammobia – Deshayes, 1855c: 319. — Type locality: New Zealand; Cuming coll. — Type age: Recent. — Type material: MNHN-IM-2000-35993, lectotype ( Willan 1993); MNHN- IM-2000-35994, paralectotype. — Current status: synonym of Gari ( Gobraus) strangeri (J. E. Gray, 1843) [ Psammobia ] ( Powell 1979: 418; Willan 1993: 49-50, fig. 207 – lectotype).

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Psammotella – H. Adams & A. Adams, 1856 (Adams & Adams 1856: 393), ex Deshayes ms, non Herrmannsen, 1852. — Type species (M): Solen ruber Schroter, 1788 . — Current status: synonym of Hiatula Modeer, 1793 .

REMARK

Deshayes apparently added manuscript names on labels in Cuming’s material of this family, but he never published the species involved. Reeve then credited Deshayes with the following ms names, although Reeve must be considered their author. These Reeve species are listed here along with one other species credited as “ ex Deshayes ms”.

acuminata View in CoL , Soletellina – Reeve, 1857a: pl. 3, fig. 12, ex Deshayes ms. Soletellina acuminata Reeve, 1857 ( Willan 1993: 75). — Current status: synonym of Hiatula diphos (Linnaeus, 1771) View in CoL [ Solen View in CoL ] ( Huber, 2015: chapter 5 on CD), the type species of Hiatula Modeer, 1793 View in CoL .

adamsii View in CoL , Soletellina – Reeve, 1857a: pl. 2, fig. 5, ex Deshayes ms. — Current status: Soletellina adamsii (Reeve, 1857) ( Willan 1993: 75) .

ambigua , Psammotella View in CoL – Reeve, 1857b: pl. 1, fig. 5, ex Deshayes ms. — Current status: Soletellina ambigua Reeve, 1857 ( Willan 1993: 67), now Hiatula ambigua (Reeve, 1857) View in CoL .

appendiculatus View in CoL , Solen ( Macropsammus) – Listed by Cossmann & Peyrot (1910: 295, fig. 44) as a Deshayes species, it was actually made available by Lamarck, 1806. Lamarck’s species is the type species (OD) of Psammoica Dall, 1900, non Solier, 1835 [ Coleoptera ], which was renamed Macropsammus Cossmann, 1902 View in CoL . — Current status: Macropsammus appendiculatus ( Lamarck, 1806) View in CoL .

atrata , Soletellina – Reeve, 1857a: pl. 3, fig. 14, ex Deshayes ms. Soletellina atrata Reeve, 1857 ( Willan 1993: 75; Huber 2010a: 330). Matsukuma, in Okutani (2017: 1262), attributed this species to “Deshayes in Reeve”. — Current status: Hiatula atrata (Reeve, 1857) View in CoL .

cumingiana , Soletellina – Reeve, 1857a: pl. 1, fig. 4, ex Deshayes ms. Soletellina cumingiana Reeve, 1857 ( Willan 1993: 75). — Current status: synonym of Hiatula adamsii (Reeve, 1857) View in CoL ( Huber 2015: chapter 5 on CD).

epidermia View in CoL , Soletellina – Reeve, 1857a: pl. 1, fig. 3, ex Deshayes ms. — Current status: synonym of Soletellina biradiata (W. Wood, 1815) [ Solen View in CoL ] ( Willan 1993: 77), now Hiatula biradiata (W. Wood, 1815) View in CoL .

incerta View in CoL , Soletellina – Reeve, 1857a: pl. 3, fig. 13, ex Deshayes ms. — Current status: synonym of Soletellina alba ( Lamarck, 1818) [ Psammobia ] ( Willan 1993: 79), now Hiatula alba ( Lamarck, 1818) View in CoL .

japonica View in CoL , Soletellina – Reeve, 1857a: pl. 4, fig. 16, ex Deshayes ms. — Current status: Nuttallia japonica (Reeve, 1857) ( Huber 2010a: 331) View in CoL . Kantor & Sysoev (2005: 362), as “Deshayes in Reeve”.

malaccensis View in CoL , Psammotella View in CoL – Reeve, 1857b: pl. 1, fig. 2, ex Deshayes ms. — Current status: synonym of Hiatula ambigua (Reeve, 1857) ( Willan 1993: 67) View in CoL .

nymphalis, Soletellina – Reeve, 1857a: pl. 1, fig. 2, ex Deshayes ms. — Current status: synonym of Soletellina biradiata (W. Wood, 1815) [ Solen View in CoL ] ( Willan 1993: 77), now Hiatula biradiata (W. Wood, 1815) View in CoL .

oblonga , Psammotella View in CoL – Reeve, 1857b: pl. 1, fig. 7, ex Deshayes ms. — Current status: synonym of Hiatula rosea (Gmelin, 1791) View in CoL [ Solen View in CoL ] ( Huber 2015: chapter 5 on CD).

obscurata View in CoL , Soletellina – Reeve, 1857a: pl. 4, fig. 21, ex Deshayes ms. — Current status: Nuttallia obscurata (Reeve, 1857) ( Huber 2010a: 331) View in CoL . Kantor & Sysoev (2005: 362) cited this as “Deshayes in Reeve”.

ovalina View in CoL , Donax View in CoL – Reeve, 1854d: pl. 3, fig. 17, ex Deshayes ms. No locality indicated.Deshayes, 1855c: 352. Central America; Cuming coll. — Current status: Synonym of Heterodonax pacificus (Conrad, 1837) View in CoL [ Psammobia ] ( Coan & Valentich-Scott 2012: 689).

philippinensis , Psammotella View in CoL – Reeve, 1857b: pl. 1, fig. 1, ex Deshayes ms. — Current status: synonym of Hiatula ambigua (Reeve, 1857) ( Willan 1993: 67) View in CoL .

subradiata , Psammotella View in CoL – Reeve, 1857b: pl. 1, fig. 6, ex Deshayes ms. — Current status: synonym of Hiatula ambigua (Reeve, 1857) ( Willan 1993: 67) View in CoL .

tellinaeformis, Psammobia – Reeve, 1856: pl. 5, fig. 31, ex Deshayes ms. — Current status: synonym of Gari livida ( Lamarck, 1818) View in CoL [ Psammobia ] ( Willan 1993: 41).

tumens View in CoL , Soletellina – Reeve, 1857a: pl. 4, fig. 20, ex Deshayes ms. — Current status: Soletellina tumens Reeve, 1857 ( Willan 1993: 89), now Hiatula tumens (Reeve, 1857) View in CoL .