Cochlis pseudoprotracta, Harzhauser & Landau & Guzhov, 2025
publication ID |
https://doi.org/10.11646/zootaxa.5703.1.1 |
publication LSID |
lsid:zoobank.org:pub:78B0FE76-1698-4FA0-99B3-661DBB27DFF6 |
persistent identifier |
https://treatment.plazi.org/id/03A687A6-6015-FFE6-FF00-F90EFCFBF99F |
treatment provided by |
Plazi |
scientific name |
Cochlis pseudoprotracta |
status |
sp. nov. |
Cochlis pseudoprotracta nov. sp.
Figs 2E View FIGURE 2 , 16A–D View FIGURE 16
Natica epiglottina Lam.—Pusch 1836: 543 View in CoL [ non Natica epiglottina Lamarck, 1804 View in CoL ].
Natica epiglottina Lam.—Pusch 1837: 99 View in CoL , pl. 9, fig. 12 [ non Natica epiglottina Lamarck, 1804 View in CoL ].
Natica ( Natica) neglecta Mayer-Eymar View in CoL — Hilber 1893: 1029, figs 8–9 [ non Cochlis neglecta (Mayer-Eymar, 1858) View in CoL ].
Natica millepunctata Lam. View in CoL var.— Friedberg 1923: 426, pl. 26, fig. 3, text-fig. 76 [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].
Natica View in CoL ( N [ atica].) aff. epiglottina Lam.— Báldi 1960: 64 View in CoL , pl. 2, fig. 2b [ non Natica epiglottina Lamarck, 1804 View in CoL ].
Natica ( Nacca) millepunctata var. tigrina ( Defrance 1825) View in CoL —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 118, pl. 33, fig. 4 [ non Cochlis tigrina ( Defrance, 1825) View in CoL ].
Natica millepunctata Lamarck, 1822 View in CoL — Zelinskaya et al. 1968: 175, pl. 41, fig. 19 [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].
Natica tigrina Röding, 1798 View in CoL — Bałuk 1995: 194, pl. 15, figs 10, 12, 14 [ non Cochlis tigrina ( Defrance, 1825) View in CoL ].
Natica tigrina ( Defrance, 1825) View in CoL — Mikuž 2009: 18, pl. 4, figs 52–53 [ non Cochlis tigrina ( Defrance, 1825) View in CoL ].
Type material. Holotype: NHMW 2024 View Materials /0232/0002 (A 1260), SL: 17.0 mm, MD: 15.2 mm, Bad Vöslau ( Austria), Figs 16C View FIGURE 16 1 –C View FIGURE 1 4 View FIGURE 4 . Paratypes: NHMW 2024 View Materials /0236/0002, SL: 16.8 mm, MD: 15.3 mm, Bad Vöslau ( Austria), Figs 2E View FIGURE 2 , 16A View FIGURE 16 1 –A View FIGURE 1 4 View FIGURE 4 . NHMW 2024 View Materials /0236/0001, SL: 16.9 mm, MD: 16.5 mm, Bad Vöslau ( Austria), Figs 16B View FIGURE 16 1 –B View FIGURE 1 4 View FIGURE 4 . NHMW 2024 View Materials /0237/0001, SL: 14.7 mm, MD: 13.9 mm, Jaroměřice ( Czech Republic), Figs 16D View FIGURE 16 1 –D View FIGURE 1 4 View FIGURE 4 .
Additional paratypes. NHMW 2024 View Materials /0236/0003, SL: 17.9 mm, MD: 17.3 mm, Bad Vöslau ( Austria). BkKG460-464 , SL: 27.0 mm, MD: 22.0 mm, Korytnica ( Poland), illustrated in Bałuk (1995: pl. 15, fig. 12) . BkK-G460- 464 , SL: 17.8 mm, MD: 16.5 mm, Korytnica ( Poland), illustrated in Bałuk (1995: pl. 15, fig. 10) .
Type locality. Bad Vöslau ( Austria), Vienna Basin.
Type stratum. Baden Formation.
Age. Middle Miocene, middle Badenian (Langhian).
Etymology. Referring to the similarity with Euspira protracta ( Eichwald, 1830) .
Diagnosis. Small, ovate shell with moderately high spire of strongly convex whorls; moderately wide umbilicus with narrow, strongly raised funicle, and small semicircular umbilical callus.
Description. Small, ovate shell of about 4.5 teleoconch whorls; higher than wide (SL/MD = ~1.1). Spire slightly elevated, broad turbiniform, of strongly convex whorls; apical angle 100–117°. Protoconch of about 2.5 convex whorls. Suture linear, weakly impressed. Faint, low subsutural cord. Narrow, weakly concave subsutural ramp, delimited by rounded shoulder. Last teleoconch whorl weakly expanding, not inflated, attaining ~90% of total height. Some specimens with very faint, widely spaced spiral cords on last whorl. Periphery strongly convex, placed slightly below mid-whorl (~49%). Growth lines very prominent, fold-like, prosocline in apical view, fading at shoulder, delicate to subobsolete at periphery. Aperture moderately wide, ovate D-shaped, with maximum diameter in abapical third. Position of adapical tip of aperture moderately high (~82%), far below shoulder. Aperture attaining ~70% of total height. Columellar lip straight, weakly reinforced, basal lip weakly reinforced, outer lip thin. Columellar angle 26–38°. Opercular ridge in outer lip weak to subobsolete. Umbilicus moderately wide (~17%), deep, semicircular. Parietal callus moderately long (~43%), weak, narrow with very small, subtrigonal anterior lobe. Margin of parietal callus weakly concave. Very narrow, raised, cord-like funicle terminating in abapical third of columellar lip in small or insignificant semicircular umbilical callus. Adapical and abapical sulci moderately wide, weakly concave, abapical sulcus delimited by faint spiral ridge. Color pattern of numerous, small, circular dots ( Bałuk 1995: pl. 15, fig. 10). Operculum unknown.
Discussion. This species has frequently been reported in the literature but was confused with the extant Mediterranean Naticarius stercusmuscarum ( Gmelin, 1791) and the Early Miocene northeastern Atlantic Cochlis tigrina ( Defrance, 1825) . It differs from both species and from most Paratethyan Cochlis species, except for Cochlis neglecta ( Mayer, 1858) , in its high spire and ovate outline. In addition, it is characterized by its narrow but strongly raised funicle. Due to its ovate outline, this species is reminiscent of Euspira protracta ( Eichwald, 1830) but differs clearly from that species in its prominent funicle. Cochlis neglecta ( Mayer, 1858) is almost identical in outline but differs in its more prominent, pillar-like funicle and lacks the characteristic, fold-like growth lines on the subsutural ramp ( e.g., Robba et al. 2016: pl. 2, fig. 14). In addition, Cochlis pseudoprotracta nov. sp. was a deep water species whereas C. neglecta is found in shallow water deposits.
Paleoenvironment. Occurrences in the Baden Formation of the Vienna Basin suggest middle to outer neritic settings in up to 250 m water depth ( Kranner et al. 2021) .
Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1995); Vienna Basin: Bad Vöslau, Baden, Baden-Sooss ( Austria) ( hoc opus); Pannonian Basin: Szokolya ( Hungary) ( Báldi 1960); Krka Basin: Ivanji dol, Šentjernej ( Slovenia) ( Hilber 1893; Mikuž 2009); Dacian Basin: Dobrusha ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Genus |
Cochlis pseudoprotracta
Harzhauser, Mathias, Landau, Bernard M. & Guzhov, Aleksandr 2025 |
Natica tigrina ( Defrance, 1825 )
Mikuz, V. 2009: 18 |
Natica tigrina Röding, 1798
Baluk, W. 1995: 194 |
Natica millepunctata
Zelinskaya, V. A. & Kulichenko, V. G. & Makarenko, D. E. & Sorochan, E. A. 1968: 175 |
Natica
Baldi, T. 1960: 64 |
Natica ( Nacca ) millepunctata var. tigrina ( Defrance 1825 )
Kojumdgieva, E. & Strachimirov, B. 1960: 118 |
Natica millepunctata Lam.
Friedberg, W. 1923: 426 |
Natica ( Natica ) neglecta
Hilber, V. 1893: 1029 |