Cylindriscala rugata, Harzhauser & Landau, 2025

Cylindriscala rugata sp. nov.

Figs 20H View FIGURE 20 1 –H View FIGURE 1 2 View FIGURE 2 , 21F View FIGURE 21

Type material. Holotype, NHMW 1871 View Materials /0010/0128, SL: 3.8 mm, MD: 1.4 mm, Porzteich at Břeclav ( Czech Republic), Figs 20H View FIGURE 20 1 –H View FIGURE 1 2 View FIGURE 2 , 21F View FIGURE 21 .

Type locality. Porzteich at Břeclav ( Czech Republic), Vienna Basin .

Type stratum. Baden Formation.

Age. Middle Miocene, middle Badenian (Langhian).

Etymology. Rugatus (Latin for wrinkled), referring to the broad, fold-like axial ribs.

Diagnosis. Very small, slender shell with strongly convex, faintly angled whorls, deeply impressed suture, robust, convex axial ribs crossed by six broad, faint spiral cords, marked basal angulation, prominent peribasal cord.

Description. Shell very small, slender (apical angle ~18°). Protoconch and first teleoconch whorls unknown. Teleoconch of at least eight strongly convex whorls with indistinct, convex subsutural shelf and faint shoulder angulation at periphery placed just above mid-whorl. Suture deeply impressed, linear. Sculpture of moderately close-set, robust, moderately raised, weakly prosocline, convex axial ribs, not aligned axially (11: 11: 11), adapical tips of ribs fading at suture. Interspaces between ribs slightly wider, with about six very weak, broad, low spiral cords, weakly overrunning axial ribs. Last whorl attaining about 37% of total height, with distinct basal angulation delimited by moderately prominent peribasal cord, basal disc flattened, nearly smooth. Aperture ovate, columella weakly concave. Outer lip weakly thickened. Columellar callus forming narrow rim, detached from base, parietal callus delicate. Peristome simple. Umbilicus reduced to narrow chink.

Discussion. This species is reminiscent of Cylindriscala bellissima sp. nov. in its angled whorls but is broader, has relatively higher whorls and the axial ribs are wider but less elevated. Furthermore, C. bellissima lacks spiral sculpture. The extant Cylindriscala acus ( Watson, 1883) is higher and has more numerous and narrower axial ribs (see Bouchet & Warén 1986: fig 1236–1237; Weil et al. 1999: fig. 82).

Paleoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Porzteich at Břeclav ( Czech Republic) (hoc opus).

Genus Discoscala Sacco, 1890

Type species. Scalaria scaberrima Michelotti, 1840 View in CoL ; original designation by Sacco (1890: 320). Miocene, Italy.

Diagnosis. “ Thin shell. Medium size ; shape turriculate, conical; whorls very convex, not disjointed, but gradate with ramp; at the anterior part of each whorl, there is a depression that is limited above by a bead invisible under the following whorl; ornamentation composed of axial, straight lamellae, deviated on the posterior ramp, not corresponding from one whorl to the next, crenellated by large spiral cords. Last whorl equal to or greater than a third of the total height, with flattened base covered by a decussate disc, limited at the periphery by a crenellated rim; the lamellae cross it and extend, a little sinuously, as far as the disc; no umbilicus in the center of the disc. Aperture rounded, subangular in front, auricle resembling a slightly calloused beak; peristome barely thickened, almost discontinuous, the inner, varnished layer becoming much thinner in the parietal region.” (translated from Cossmann 1912: 59).

Discussion. Discoscala is reminiscent of Cirsotrema , but its axial lamellae do not have a tendency to fuse, its lamellae stop abruptly at the peribasal cord and the peribasal cord delimits a well-developed basal disc that bears fine spiral threads and no axial sculpture except for growth lines.