Empis (Enoplempis) mixopolia Melander

Empis (Enoplempis) mixopolia Melander View in CoL

( Figs 268–270 View FIGURES 266–274 , 276 View FIGURES 275–276 , 284 View FIGURES 284–285 )

Empis mixopolia Melander, 1902: 327 View in CoL . Type locality (by lectotype designation): Kendrick , Idaho ( USA).

E. (Enoplempis) mixopolia View in CoL : Melander, 1928: 142 (assigned to subgenus Enoplempis ).

Empis mixopolia View in CoL : Melander, 1965: 460 (catalogue); Yang et al. (2007): 137 (catalogue).

Type material examined. LECTOTYPE (here designated in order to fix identity of the species) ♂, labelled: “ Kendrick [46°36′N 116°39′W]/ Idaho ”; “ JM Aldrich/ Coll”; “Cotype/ No. 41823/ U.S.N.M. [red label]”; “ LECTOTYPE / Empis mixopolia Melander / des. Sinclair, Brooks &/ Cumming 2025 [red label]” ( USNM) GoogleMaps . PARALECTOTYPES: USA. Idaho: Same data as lectotype (2♂, AMNH; 7♂, 1♀, USNM). Latah County: Juliaetta [46°34′N 116°42′W], J.M. Aldrich (1♂, 1♀, USNM) GoogleMaps ; Moscow [46°48ʹN 116°52ʹW] (2♀, AMNH; 3♂, USNM) ; Troy [46°44′N 116°46′W], 4.v.1899, J.M. Aldrich (1♂, 1♀, USNM) GoogleMaps .

Notes on type specimens. This species was described from an unspecified number of male and female specimens collected by J.M. Aldrich deposited in the AMNH and USNM. Two male and two female paralectotypes are also present in OUMNH.

Additional material examined. USA. Idaho: Latah County: 3 mi. S Helmer [46°48ʹN 116°28ʹW], 22.v.1971, WJT (2♂, 1♀, WFBM) ; Juliaetta [46°34′N 116°42′W] (1♀, USNM); Kendrick [46°36′N 116°39′W] (2♀, USNM); Moscow [46°48ʹN 116°52ʹW] (1♀, USNM); same data except, 10.v.1936, 2560 ft, R GoogleMaps . E. Miller (1♂, WFBM); Moscow Mtn [46°48ʹN 116°52ʹW], 17.vi.1904 (1♀, USNM); same data except, 26.v.1918, ALM (1♂, USNM) GoogleMaps ; Paradise Ridge, 46°39′36.7″N 117°22′19.3″W, 24.v.2022, A. Hawse, ex Lomatium multiflorum (1♂, CNC); same data except, 24.v.2022, ex Lomatium triternatus (1♂, 1♀, CNC); same data except, 31.v.2022, ex Lithophragma parviflorum (1♀, CNC); same data except, 31.v.2022, ex Lomatium triternatus (1♀, CNC); Troy , Hwy 8 at mm [milemarker] 15, West Fk Ck , 747 m, 46.733°N 116.757°W, 9.vi.2009, E09-62b, I.S. Winkler (1♂, CNC). Nez Perce County: Craig Mts [46°09′N 116°44′W], 8.vi.1918, ALM (1♂, USNM) GoogleMaps ; Lk. Waha [46°12ʹN 116°50ʹW], 9.vi.1918, ALM (1♀, USNM) . Owyhee County: Givens Springs [43°24′N 116°42′W], 3 mi. E, Stanleya pinnata , 30.v.1962, A. R GoogleMaps . Gittens (1♂, 3♀, WFBM). Washington County : 2 mi. N Mann’s Ck [44°26′N 116°55′W], 31.v.1962, A. R GoogleMaps . Gittens (1♂, WFBM) . Oregon: Baker County: L. Goose Ck, 36 mi. SE Union [44°56′N 117°25′W], Wallowa-Whitman NF GoogleMaps , 4600 ft, 9.vi.1977, WJT (1♂, WSU) ; U. Goose Ck , 34 mi. SE Union [44°56′N 117°25′W], 4160 ft, 24–25.vi.1977, sweeping Anaphalis flowers, WJT (13♂, 4♀, WSU) GoogleMaps . Umatilla County: 10 mi. E Weston [45°47′N 118°15′W], 24.v.1969, yarrow bloom, K. Goeden (1♂, 2♀, ODAC). Union County : Jordan Ck, 28 mi. SSW La Grande , 4840 ft, Wallowa-Whitman NF GoogleMaps , 16.vi.1977, WJT (1♂, 1♀, WSU) . Wallowa County: Minam SP [45°47′N 117°43′W], 9.vi.1971, WJT (5♂, 2♀, WFBM) GoogleMaps . Washington: Asotin County: 5 mi. S Anatone, Fields Spring SP [46°4′N 117°10′W], 17.vi.1984, J. Jenkins (7♂, 5♀, WSU); same data except, J.M. Sirota (1♂, 2♀, WSU); Fields Spring SP, 31.vii.1971, MT GoogleMaps dry ice, WJT, W.B Garnett (1♂, WSU); same data except, 14.vi.1972, dry ice MT GoogleMaps trap, WJT (8♂, 1♀, WSU) ; same data except, 15.vi.1972 (4♂, 3♀, WSU); same data except, 26.vi.1972 (1♂, 1♀, WSU); same data except, 27.vi.1972 (1♂, WSU); Fields Spring SP, 4 mi. S Anatone , 3500–4000 ft, 7.vi.1973, WJT (2♂, 2♀, WSU) ; same data except, D. Corredor (2♂, 2♀, WSU); same data except, M. Jackson (1♂, WSU); same data except, 12–13.vi.1974, MT with dry ice, WJT (10♂, 6♀, WFBM) ; same data except, 26.v.1974 (1♂, WSU); same data except, ex: flowers Taraxicum [sic] officinale (1♂, 1♀, USNM; 2♂, 16♀, WSU). Columbia County: Tucannon Fish Hatch. , 20 mi. E Dayton [46°19′N 117°39′W], 2650 ft, 15.v.1979, MT, WJT (5♂, WSU) GoogleMaps ; same data except, 12.v.1979, sweeping (1♀, WSU). Whitman County: Almota [46°42′N 117°28′W], 25.v.1913 (1♂, USNM); same data except, 6.v.1972, dry ice MT, WJT (11♂, 5♀, WSU) GoogleMaps ; same data except, 27.vi.1972 (2♂, WSU); Colton [46°34′N 117°7′W], 24.v.1952, S.E. Knapp (1♂, USNM); Kamiak Butte, 46°53′02.8″N 117°09′04.4″W, 20.v.2022, A. Hawse, ex Lomatium triternatus (1♂); same data except, 11.v.2023 (1♀, CNC); same data except, Lomatium multiflorum (2♀, CNC); Kramer Strip Nat. Palouse , 7 mi. NW Colton [46°36′N 117°14′W], 16.v.1987, sweep, WJT (1♂ WSU) GoogleMaps ; Palouse Nat.Area [46.73°N 117.16°W], 5 mi. N Colton, 16.v.1987, J.B. Johnson (1♀, WFBM); Pullman nr SCS Pond-WSU [46°43′N 117°9′W], 29.v.1971, WJT (1♀, WSU) GoogleMaps ; Pullman [46°43′N 117°10′W], 19.v.1912 (1♂, 2♀, USNM); same data except, 30.v.1913 (1♀, USNM); same data except, 10.vi.1910 (1♀, USNM); same data except, 20.v.1917, ALM GoogleMaps ; same data except, 6.vi.1918 (1♂, USNM); 8 mi. SW Pullman, Lyle Grove Biol. Area [46°43ʹN 117°11ʹW], 2100 ft, 26.v.1979, sweep, WJT (1♀, WSU) GoogleMaps ; same data except, 1.vi.1979, MT GoogleMaps , S. Woods (2♀, WSU); same data except, 4.vi.1979 (2♂, WSU); same data except, 4–6.vi.1979 (1♂, WSU); same data except, 8– 13.vi.1979 (1♂, WSU); Upper Union Flat Rd, 46°46′47.0″N 117°22′19.3″W, 28.v.2022, A. Hawse, ex Balsamorhiza sagittata (1♂, CNC); Wawawai [46°38′N 117°22′W], 2400 ft, 28.v.1922, ALM (2♂, CNC; 2♂, 2♀, USNM) GoogleMaps ; 2 mi. E Wawawai [46°38′N 117°22′W], 7.v.1965, L.S. Hawkins, Jr (1♂, WFBM) GoogleMaps .

Diagnosis. This species is distinguished from other species of the E. (En.) valentis species group by the occiput bearing two irregular rows of strong setae ( Fig. 268 View FIGURES 266–274 ), four marginal scutellar setae, tuft of black basal costal setae distinct from distal costal setae, male hind femur with tuft of setae arising directly from inflated anteroventral process and without additional lobes ( Figs 269, 270 View FIGURES 266–274 ), and wing length usually less than 7 mm.

Redescription. Wing length 5.8–7.0 mm. Male. Similar to E. (En.) aeripes , except as follows: head with occiput bearing two irregular rows of strong setae ( Fig. 268 View FIGURES 266–274 ).

Thorax with scutum with 2 pairs of broad brownish vittae, generally fading beyond transverse suture. Scutum with 1–2 posterior npl. Scutellum with 2 pairs of strong sctl, widely separated medially.

Legs with coxae clothed in dense greyish pruinescence, concolourous with pleura dorsally changing to orange brown ventrally, similar to femora. Hind femur and tibia with similar modified processes and setae as E. (En.) aeripes ( Figs 268–270 View FIGURES 266–274 ).

Wing with tuft of black basal costal setae distinct from row of distal costal setae.

Terminalia ( Fig. 276 View FIGURES 275–276 ): indistinguishable from E. (En.) aeripes .

Female. Similar to male, except as follows: hindleg without modified setae and processes; abdominal pleural membrane with darkened, expanded region.

Geographical distribution and seasonal occurrence ( Fig. 284 View FIGURES 284–285 ). Empis (En.) mixopolia is known from eastern Washington, eastern Oregon, and Idaho. Adults have been collected from May to July.

Nuptial gift presentation. Form unknown.