Exphora angustivenosa Junkiert & Walczak, 2021

Exphora angustivenosa Junkiert & Walczak View in CoL sp. nov.

( Figs 1C View Fig , 2C View Fig , 3C View Fig , 10–11 View Fig View Fig )

urn:lsid:zoobank.org:act:

Material examined

Holotype

1 ♂ / Exphora angustivenosa sp. nov. / Junkiert & Walczak det. 2018 [red label]; Madagascar / Nosy Komba [blue label]; Coll. R.I.Sc.N.B. [blue label] ( RBINS).

Paratypes

1 ♂ / Exphora angustivenosa sp. nov. / Junkiert & Walczak det. 2018 [red label]; Coll. R.I.Sc.N.B. / Madagascar / Nossi-Be / Ex Coll. Le Moult [green label] ( RBINS) ;

1 ♀ / Exphora angustivenosa sp. nov. / Junkiert & Walczak det. 2018 [red label]; Coll. R.I.Sc.N.B. / Madagascar / Nosy Komba [blue label] ( RBINS) .

1 ♀ / Exphora angustivenosa sp. nov. / Junkiert & Walczak det. 2018 [red label]; Coll. R.I.Sc.N.B. [blue label]; INSTITUT / SCIENTIFIQUE / MADAGASCAR [white label]; Madagascar Sambirano / Nosy-Be / forêt de Lokobe / I-60 / Andria Robinson [white label] ( RBINS) .

Description

Body length 12.35–12.60 mm.

Head. Metope twice as long as wide, lateral margins slightly arched and extended laterally at level of 3/ 4 in its lower part, then narrowing to metopoclypeal suture. Dorsal margin of metope weakly arcuately but distinctly concave. Median keel distinct, running through metope and metopoclypeal suture ( Fig. 2C View Fig ). Metopoclypeal suture elongate (about 1/4 shorter than metope), triangular. Lateral keels present and distinct. In lateral view metope is distinctly convex, whereas metopoclypeal suture almost straight. Median keel in lateral view gently arches above the surface of the metopoclypeal suture and it also protrudes clearly above surface of metope, so that in lateral view it is still visible, and rather not covered by slightly protruding lateral keels. The eyes not quite round (hind edge of the eye almost straight). Coryphe almost 1.5 times as wide as long, with anterior margin strongly convex and strongly angular, and posterior margin distinctly and very arcuately concave.

Pronotum and mesonotum. Pronotum bluntly rounded anteriorly, distinctly concave posteriorly. Mesonotum weakly convex, with three parallel keels: median keel is in its anterior part joined to two slanting lines converging to each other almost at right angle and thus forming an arrow-shaped structure. Both edges of arrow are joined to two lateral keels.

Fore wings. Clavus elongate, as long as 2/3 of whole wing length, of hyperpterism type. Costal area well developed with 10 or 11 cells between CA and Pc+CP; ScP+R short, separating; RA two-branched; RP with 5–7 terminals; MP separating before nodal line; MP1+2 separating before nodal line, after the third or fourth terminal, clearly forked at the same level as stigma; MP3+4 separating, before nodal line; MP4 single, MP3 short, separating before nodal line, after the first terminal. CuA forking before nodal line and before MP. Two or three transverse veins connecting CuP and PCu+A1 (one vein connecting CuP and PCu before joining with A1 and one or two connecting CuP and PCu+A1). A1 running parallel to posterior margin of clavus. Cubital cell at least twice as long as postcostal cell, radial cell and median cell. C1 based well before, C2, C3 and C4; C5 starts at a similar level as C1; C2 and C 4 in contact sharing MP margin. C1, C2, C3 and C4 of similar length, C5 about 2 times longer. Eighteen to nineteen apical cells. Stigma longitudinal and brown, well visible, includes 3-4 (mostly 3) short veins running to edge of wing ( Fig. 3C View Fig ).

Hind wings: Well developed, 4/5 of the fore wings’ length. Hyaline with brown veins. Almost twice long as wide in midline (ratio length to width: 2.2–2.4). Eleven apical cells ( Fig. 3C View Fig ).

Legs. Prothoracic and mesothoracic legs: Femur about 2/3 of tibia length, irregular in cross-section, margins of femur covered with small bristles, inner margin bear small spines. Tibiae long and thin, trapezoidal in cross-section, margins covered with small bristles, lateral margins with double row of bristles.

Metathoracic legs: Metafemur margins covered with small bristles. Metatibia twice as long as metafemur, triangular in cross-section with concave ventral side. Margins of metatibia covered with small, barely visible bristles. Lateral margin with four lateral spines, three of which are distinct and one is weakly visible. Metatibiotarsal formula 8/8/2.

Colouration. The species is paler than the previous one. General colouration ochre-lightbrown, metope with keels distinctly red, between lateral and median keels red-brown thin stripes passing (much thinner than the previous two species) passing along the metope. Eyes darker than background, almost black, ocelli with delicate rim, blurred at the back. Posterior margin of pronotum and mesonotum gently contoured by a very thin dark line, all convex keels on the mesonotum also darker than the background. Mesonotum at the proximal part with two small spots, with two larger irregular spots on each side, and at the distal part with two small but distinct spots. Lateral margins of tergites of abdomen light brown. Wings hyaline with red-brown veins. Legs ochre-lightbrown, metatibial spines of the same colour except for the tip that is black.

Genitalia. Male. Pygofer with hind margin convex. Aedeagus s.l. arched, robust in lateral view. Ventral phallobase not reaching half-length of aedeagus. Each dorso-lateral phallobase lobe with long fusiform process reaching the apex of aedeagus, apical process semi-circularly covered by abundant denticles, the lateral margin serrate, subapical process absent. Inner phallobase lobes spatulate, slightly curved, narrow on the base and wider on apices. Aedeagus s.s. long and fusiform ( Fig. 10A View Fig ). Anal tube rather robust, enlarged apically in dorsal view with concave lateral margin and roundish lateral margins. Anal column length about 0.54 times anal tube length ( Fig. 10B View Fig ). Gonostyle oval, elongated. Capitulum wide and folded, bearing subapical tooth on inner side ( Fig. 10C View Fig ).

Female. Endogonocoxal process, well developed, apical ends roundish, with well visible incision, inner margin strongly sclerotized. Gonoplac (third valvula) semi-circular in lateral view ( Fig. 11A View Fig ). Median part of sternite VII with distinct roundish incision in the middle. Genitalia with anal tube relatively short, broadly rounded in dorsal view ( Fig. 11B View Fig ). Gonapophyses VIII (first valvula) almost trapezoidal, elongated, strongly sclerotized and bearing three teeth on dorsal margin and one large, long tooth, bearing smaller subapical tooth ( Fig. 11C View Fig ). Gonapophyses VIII not visible externally, covered by gonoplacs. Gonapophyses IX (second valvular) sclerotized, confluent with endogonocoxal processes ( Fig. 11D View Fig ).

Measurements (in mm, females in parentheses). Body length: 12.35 (12.60); head width (with eyes): 2.00 (1.93–2.00); metope length: 2.18 (1.84–2.14); metope width: 1.28 (1.27–1.30); metope length/metope width ratio: 1.70 (1.41–1.68); coryphe length: 0.68 (0.60– 0.67); coryphe width: 0.98 (0.83–0.90); coryphe length/ coryphe width ratio: 0.69 (0.72–0.74); fore wing length: 10.46 (10.17–10.99); fore wing width: 5.13 (4.78–5.32); mesonotum length: 2.40 (2.32–2.36); mesonotum width: 2.25 (2.12–2.15).

Diagnosis. Exphora angustivenosa sp. nov. can be distinguished by its size (it is bigger than other representatives of this genus), small brown longitudinal stigma which includes 3–4 (mostly 3) short veins running to edge of wing, metope with thin brown stripes, anal tube with roundish lateral margins, shape of style, aedeagus and its processes.

Etymology. The name of species refers to the thin veins on the fore wings.

Distribution. Madagascar, Nosy Ambariovato, Nosy Komba [ 13°28’18.7” S, 48°20’54.2” E]; Nosy Be, Lokobe Forest [ 13°23’36.8” S, 48°18’42.4” E] ( Fig. 7 View Fig ).