Garnotia stricta var. acutigluma, 2015

b. var. acutigluma (Steud.) Veldk. View in CoL , comb. nov.

Garnotia acutigluma (Steud.) Ohwi var. acutigluma View in CoL , automatically generated by Santos (1950: 84–85; the typical form not mentioned by name). See note. — Urachne acutigluma Steud. (1854) View in CoL 121. — Garnotia acutigluma (Steud.) Ohwi (1941) View in CoL 393. — Type: Goering II, 141 (holo P; US fragm.) (from Java, not Japan).

Streptachne indica Buse (1856) View in CoL 99. — Type: Reinwardt A º. 1818 (holo L, no. 908.97-1375), designated here.

Garnotia adscendens Munro ex Ridl. & H.J.P. Winkl. (1910) View in CoL 522 (‘ ascendens ’), nom. nud. — Voucher: Winkler 3036 (W lost?; BM, BO, L, WRSL).

Garnotia caespitosa Santos (1944) 92. — Type: BS 6237 ( Gates ) (holo US; BM, K 000290161 , NY, PNH lost).

Garnotia caespitosa Santos var. longiuscula Santos (1950) 67. — Type: BS 34069 ( Ramos & Edaño) (holo UC; A, PNH lost).

Garnotia caespitosa Santos forma permucronata Santos (1950) 67. — Type: BS 36776 ( Ramos & Edaño) (holo US; K 000943326, PNH lost).

Garnotia caespitosa Santos var. emarginata Santos (1950) 68. — Type: BS 47127 ( Ramos & Edaño) (holo US; BM, NY, PNH lost, UC).

Garnotia caespitosa Santos forma amboinensis Santos (1950) 68. — Type: BS 1649 ( Robinson ) (holo US; PNH lost).

Garnotia longiaristata Santos (1950) View in CoL 70. — Garnotia acutigluma (Steud.) Ohwi var. longiaristata Jansen (1953) View in CoL 382. — Type: Endert 2645 (holo US; BO, K, L).

Garnotia longiaristata Santos var. basilanensis Santos (1950) View in CoL 71. — Type: Ebalo 907 (holo MICH).

Garnotia erecta Santos (1950) 73. — Type: SF 13854 (Robinson) (holo K 000290154; BM, PNH lost, SING).

Garnotia erecta Santos forma malayana Santos (1950) 74. — Type: SF 23465 ( Henderson ) (holo US; SING).

Garnotia acutigluma (Steud.) Ohwi var. aberrans Santos (1950) 84. — Type: Backer 22768 (holo US; BO, L, SING, U).

Garnotia acutigluma (Steud.) Ohwi var. subvestita Santos (1950) 84. — Type: Bakhuizen v.d. Brink 3683 (holo US; BO, K, L, U).

Garnotia acutigluma (Steud.) Ohwi var. ambigua Santos (1950) 85. — Type: Backer 22401 (holo US; BO, K 000943327 , L, U, US).

Garnotia acutigluma (Steud.) Ohwi forma abbreviata Santos (1950) 85. — Type: Backer 25632 (holo US; BO, K, L, U).

Garnotia acutigluma (Steud.) Ohwi forma partiens Santos (1950) 85. — Type: Winckel 1404 (‘ 14043 ’) (holo US; BO, L, U).

Garnotia griffithii Munro (1865) View in CoL 58, nomen; Hook.f. (1896) 243, in syn. — Voucher: Griffith KD 6780 (K, L).

Garnotia stricta View in CoL auct. non Brongn.

Plants perennial. Culms erect to decumbent, branched at base to branched along the culm, 0.2–0.85 m long. Nodes glabrous to pilose. Sheath mouth glabrous to sparsely hairy. Ligule 0.1–0.5(–1.75) mm long. Blades linear, flat to folded, 3.5– 29(–40) cm by 1.7–6(–12) mm, glabrous to pilose (sparsely), base narrowly attenuate. Panicle 5–20(–35) cm long, common axis longer than the lowest branches, branches solitary to fascicled, at maturity appressed to erecto-patent, smooth to scaberulous. Spikelets 3–4.5(–5.5) mm long (excl. awn), callus hairs 0.2–0.7(–1) mm long. First glume 2.5–4.5(–5.5) mm long, shorter than to longer than the second glume, acute to mucronate. Second glume 2.5–4.25(–5) mm long, apex acute to mucronate, arista 0–1(–4) mm long. Lemma awn apical, imperfect, column absent, arista straight, 8–15 mm long. Anthers 1–1.8 mm long.

Distribution — Nepal, Sikkim, Bhutan, NE India ( Assam) to S China ( Guangdong, Guangxi, Guizhou, Yunnan), Taiwan; Thailand: Northern: Chiang Mai; North-eastern: Loei, Nakhon Phanom; South-eastern: Prachinburi; Penins.: Nakhon Sri- tamarat, Trang; Malesia: Malay Peninsula ( Kedah, Pahang, Perak), Sumatra (Padang), W Java ( Bantam, Bogor, Jakarta, Preanger), Central (Banyumas), Lesser Sunda Isl.:? Bali, Flores, Sumbawa, Borneo ( Brunei, SE Kalimantan, Sabah, Sarawak), Celebes (SW; Talaud Isl.), Philippines ( Basilan, Luzon, Mindanao), Moluccas (Ceram, Ternate). Probably introduced and naturalised in Hawai’i.

Habitat — Local, but then sometimes vegetation forming. A facultative rheophyte on boulders, sandstone rocks, also in marshy places, open grassland, evergreen forest, bamboo forest, 15–1300 m altitude.

Collector’s notes — Inflorescences pale green to green.

Notes — An autonym is generated by the description of a taxon not including the type of the specific name, even when it is not mentioned in the original text. When they are united, as here, the non-published autonym has priority (Art. 6.8, 7.6, 26.1; McNeill et al. 2012). Santos under the description of presum- ably the typical form, diagnosed the 5 varieties and formae as “similis formae typicae a qua differt…”.

The only collection seen from Sumatra ( Meijer 6038, L) is exceptional for being taller than the others, with longer leaves and spikelets.