Monacha ( Metatheba ) ciscaucasica, Hausdorf, 2000

Monacha ( Metatheba) ciscaucasica View in CoL , sp. n.

(®gures 5, 14, 19)

Helix carthusiana [ sic], Kaleniczenko 1853: 83 [ partim, non O. F. MuÈller, 1774].

Helix frequens, Mousson 1863: 318 [ partim, non Mousson, 1859].

Helix frequens, Rosen 1911: 101 [ partim, non Mousson, 1859].

Theba orientalis roseni Hesse, 1914: 263 [ partim, non Hesse, 1914].

Shell (®gure 5). Depressed conical±globular; with 4×±5× convex whorls; teleoconch with wrinkled growth-ridges and Ô distinct incised spiral striae; whitish or light corneous, then a whitish peripheral band might be discerned; body whorl rounded; aperture elliptical or almost circular; upper insertion of the peristome usually slightly descending; peristome sharp, not expanded, with a whitish internal rib; umbilicus narrow, slightly obscured by the reēcted columellar edge.

Measurements. Pyatigorsk, Mashuk Gora ( n 5 20): D: 8.3±13.9 mm, x Å 5 11.4 mm; H: 5.7±9.4 mm, x Å 5 7.6 mm; D/H: 1.43±1.58, x Å 5 1.50.

Genitalia (®gure 14). The penial papilla is Ô conical or almost cylindrical. The two glandulae mucosae are simple or little rami®ed. The bursa of the bursa copulatrix is Ô oval. The right ommatophoral retractor crosses with the penial retractor. The measurements of some sections of the genitalia are compiled in table 3 and the variation of the proportions of some sections of the genitalia is compiled in table 4.

Remarks. Monacha ciscaucasica cannot be distinguished from M. ( Paratheba) roseni by shell characters. The umbilicus of M. ciscaucasica is narrower than the umbilicus of M. subcarthusiana , but wider than the umbilicus of M. perfrequens .

The genitalia of M. ciscaucasica diOEer from the other Caucasian Metatheba species , e.g., in the length of the penial papilla which is in relation to the epiphallus longer than in M. subcarthusiana , but shorter than in M. perfrequens .

Etymology. Because of the occurrence in Ciscaucasia.

Type locality. Russia: Pyatigorsk , Mashuk Gora, LP47 .

Distribution and material (®gure 19). So far M. ciscaucasica is only known from the surroundings of Pyatigorsk in the Northern Caucasus. Only the specimens from the Mashuk Gora are paratypes.

Russia. Pyatigorsk , Mashuk Gora, LP47 (det. anat.; holotype ZMH 2771 View Materials , leg. E. Clauss, measurements: D 5 12.9 mm, H 5 8.8 mm; CLA; SMF 278702 About SMF ; ZMH 2770 View Materials ); Pyatigorsk, LP47 ( paralectotype s of roseni: SMF 6605; ZMZ 505714 ( partim?) specimens to Mousson 1863; SMF 69919, 69927 About SMF , 98770 About SMF ; ZIRAN; ZMUM 4519 View Materials , 8424 View Materials ) .

( Metatheba ) species. Abbreviations: ep5 epiphallus; ¯ 5 ¯agellum; p 5 penis; pp5

penial papilla; v 5 vagina up to the glandulae mucosae; v 5 total length of the

gm t

vagina.