Neverita manhartensis ( Schaffer, 1912 )

Neverita manhartensis ( Schaffer, 1912)

Figs 56A–B View FIGURE 56

* Natica ( Neverita) Josephinia Risso var. Manhartensis Schff. — Schaffer 1912: 166, pl. 54, figs 15–16.

N [ everita]. olla View in CoL manhartensis Schff.— Sieber 1958: 139 .

Polinices ( Neverita) olla De Serres— Hölzl 1958: 152 , pl. 9, fig. 3 [ non Neverita olla (de Serres, 1829) View in CoL ].

Polinices ( Neverita) josephinia ( Risso, 1826) — Hölzl 1958: 207, pl. 18, fig. 18 [ non Neverita josephinia Risso, 1826 View in CoL ].

Polinices ( Neverita) olla (De Serres) — Răileanu & Negulescu 1964: 175, pl. 13, fig. 10 [ non Neverita olla (de Serres, 1829) View in CoL ].

Neverita olla manhartensis Schaffer, 1912 — Steininger et al. 1971: 393, pl. 7, figs 4–5.

Polinices josephinia olla (De Serres) — Báldi 1973: 274, pl. 33, figs 2–4 [ non Neverita olla (de Serres, 1829) View in CoL ].

Natica ( Neverita) josephinia olla Serres— Nicorici & Sagatovici 1973: 170 View in CoL , pl. 25, figs 5–9 [ non Neverita olla (de Serres, 1829) View in CoL ].

? Neverita olla ( Serres 1829) View in CoL — Pfister & Wegmüller 2007: 102, pl. 18, figs 17–24, pl. 19, figs 1–7 [ non Neverita olla (de Serres, 1829) View in CoL ].

? Neverita josephinia ( Risso, 1826) View in CoL — Höltke 2009: 88, fig. 35 (unidentifiable specimen).

Type material. Lectotype (designated herein), GBA 1912 /004/18/1, SL: 20.9 mm, MD: 23.1 mm, Gauderndorf ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 15), Figs 56A View FIGURE 56 1 –A View FIGURE 1 4 View FIGURE 4 . Paralectotype, GBA 1912 /004/18/2 , SL: 14.3 mm, MD: 17.8 mm, Loibersdorf ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 6), Figs 56B View FIGURE 56 1 –B View FIGURE 1 4 View FIGURE 4 .

Revised description. Medium sized, depressed depressed-globose of about three teleoconch whorls; wider than high ( SL /MD = ~0.9). Protoconch unknown. Spire depressed, broadly conical; apical angle 140°. Suture linear, adpressed to insignificant. Teleoconch whorls with broad, moderately convex subsutural ramp; no shoulder. Last whorl strongly expanding, attaining ~90% of total height, weakly convex below subsutural ramp, moderately convex at periphery. Periphery below mid-whorl (~42%). Growth lines delicate and weakly prosocyrt in apical view, prosocline and insignificant at periphery. Aperture poorly preserved, very wide, ovately D-shaped, strongly prosocline. Position of adapical tip of aperture moderately high (~85%), distinctly above mid-whorl. Aperture attaining ~72% of total height. Columellar lip slightly concave, basal and outer lips moderately thin. Columellar angle ~13°. Opercular ridge in outer lip unknown. Umbilicus largely filled by thick, kidney-shaped umbilical callus, covering funicle. Parietal callus long (~55%), strongly thickened, slightly expanding over base, with straight margin or grading into base without clear margin. Anterior lobe of parietal callus small, slightly thickened. Broad, deep umbilical channel encircling umbilical callus. Umbilical wall with prominent growth lines. Basal fasciole broad, rounded. Color pattern not preserved.

Discussion. Neverita manhartensis ( Schaffer, 1912) is a poorly known species, documented by few poorly preserved specimens. Nevertheless, it differs from Neverita olla (de Serres, 1829) in its wide and deep umbilical channel, which justifies separation. Neverita antiqua (Saco, 1890) from the Rupelian of Italy is almost identical in outline and umbilical features and differs from N. manhartensis only in its strongly convex umbilical callus (see Sacco 1891: pl. 2, fig. 55, Robba et al. 2016: pl. 11, figs 3–4). Overall, N. manhartensis is closer to the Oligocene N. antiqua than to the Miocene N. olla . This relation might be supported by the fact, that it occurs already in the Late Oligocene of the Central Paratethys ( Báldi 1973).

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Egerian (Late Oligocene/Early Miocene): Eger, Törokbálint ( Hungary) ( Báldi 1973).

Eggenburgian (Early Miocene): North Alpine Foreland Basin: Belpberg, St. Gallen ( Switzerland) ( Pfister & Wegmüller 2007); Kaltenbachgraben, Ermingen ( Germany) ( Hölzl 1958; Höltke 2009); Gauderndorf, Mörtersdorf, Loibersdorf, Maria Dreieichen ( Austria) ( Schaffer 1912); Zârand Basin: Minişul des Sus ( Romania) ( Nicorici & Sagatovici 1973); Transylvanian Basin: Coruş ( Romania) ( Răileanu & Negulescu 1964).