Nudiflorae Hook.f. (1880)

Sect. III. Nudiflorae Hook.f. (1880) 184. —

Type: Lasianthus blumianus Wight

Key to the species

1. Cymes very shortly peduncled;branchlets conspicuously sulcate; leaves large, coriaceous, 18– 25 by 5 – 8 cm; calyx obconic-cupular, truncate or minutely 4-toothed.................................... 95. L. pergamaceus View in CoL

1. Cymes sessile or tuberculate; branchlets not sulcate; leaves various; calyx various.................... 2

2. Leaves less than 3 cm long......... 10. L. bifloroideus View in CoL

2. Leaves more than 3 cm long..................... 3

3. Leaves linear, 10–14 by 0.9–1.8 cm, long-caudate at the apex, glabrous................... 70. L. linearifolius View in CoL

3. Leaves otherwise.............................. 4

4. Leaves with conspicuously looped venation; branchlets and leaves glabrous except sometimes puberulous nerves beneath..................................... 5

4. Leaves without looped venation; branchlets and leaves glabrous or hairy.............................. 7

5. Leaves lanceolate, nerves 9–12 pairs; drupes puberulous and slightly warty............. 16. L. brochidodromus View in CoL

5. Leaves oblong or oblong-lanceolate; nerves less than 9 pairs; drupes glabrous and conspicuously warty...... 6

6. Leaves subrounded at the base; nerves 6 – 8 pairs; drupes with conspicuous 8 ridges; pyrenes with 2 furrows on the abaxial face.............. 77. L. membranaceoideus View in CoL

6. Leaves acute to cuneate at the base; nerves 3–4 pairs; drupes with 3–4 obtuse ridges; pyrenes without furrow on the abaxial face................ 63. L. kinabaluensis View in CoL

7. Flowers on pedicels 2 mm long or more............ 8

7. Flowers sessile or subsessile pedicels less than 1 mm long....................................... 13

8. Leaves thinly membranaceous; nervules weak, inconspicuous; stipules broadly ovate.. 120. L. submembranifolius View in CoL

8. Leaves chartaceous to thickly coriaceous; nervules conspicuous; stipules otherwise..................... 9

9. Leaves elliptic, oblong-elliptic or oblong, large, 4.5–8 cm broad...................................... 10

9. Leaves lanceolate, ovate-lanceolate, oblong-lanceolate, less than 4 cm broad.......................... 11

10. Leaves chartaceous or subcoriaceous with acute bases........................... 14. L. brevipedicellatus View in CoL

10. Leaves thickly coriaceous with subrounded bases.................................... 129. L. vriesianus

11. Leaves; nervules reticulate; pedicels 8 –10 mm long; drupes verrucose, c. 6 mm diam, with 4 conspicuous obtuse ridges; pyrenes 4....................... 101. L. purpureus View in CoL

11. Leaves; nervules parallel; pedicels 2– 3 mm long; drupes 3–4 mm diam, with 4–5 (smooth) or 7–8 (verrucose) obtuse ridges; pyrenes 4–5 or 7–8................. 12

12. Stipules less than 1 mm long; calyx with triangular teeth; drupes smooth, with 4–5 obtuse ridges; pyrenes 4–5................................ 92. L. pedicellatus View in CoL

12. Stipules 3–4 mm long; calyx truncate; drupes verrucose, with 7–8 obtuse ridges; pyrenes 7–8... 87. L. oblongus View in CoL

13. Leaves narrowly oblong-lanceolate, narrowly lanceolate or linear-lanceolate, 5 – 6 times longer than broad, longer than 12 cm ................................. 14

13. Leaves otherwise............................ 17

14. Plants densely tomentose; leaves 23–25 by 3–4 cm; nerves 16–18 pairs; calyx with 7–8 lanceolate lobes.......................................... 41. L. eriocalyx View in CoL

14. Branchlets glabrous or pubescent; leaves 9–19 by 1.5–3 cm; nerves less than 15 pairs; calyx with 4–5 teeth or with an almost truncate limb........................ 15

15. Nerves 7 pairs; stipules narrowly triangular, c. 4 mm long; calyx almost truncate or minutely 5-toothed...................................... 1. L. acuminatissimus View in CoL

15. Nerves 8–14 pairs; stipules triangular, 2–3 mm long; calyx with 4–5 conspicuous teeth.................... 16

16. Branchlets glabrous; leaves with densely hirsute margin; nerves 8 –11; petioles 12–15 mm long.......................................... 57. L. hirtimarginatus View in CoL

16. Branchlets densely appressed-pubescent; leaves without hirsute margin; nerves 12–14 pairs; petioles 4–5 mm long............................... 3. L. angustifolius View in CoL

17. Drupes usually with 2 pyrenes (only in Borneo)..... 18

17. Drupes with 3 or more pyrenes.................. 20

18. Leaves membranaceous; drupes ovoid, very rugose, without cork-like structure between pyrenes; pyrenes rugose on the abaxial face........... 78. L. membranaceus View in CoL

18. Leaves coriaceous; drupes ellipsoid, smooth, with cork-like structure between pyrenes; pyrenes smooth on the abaxial face....................................... 19

19. Plants glabrous; leaves with subrounded base, nervules conspicuously prominent on both surfaces; petioles 5 –7 mm long; stipules c. 2 mm long; calyx tube 3 mm long, not expanded at limb................. 84. L. nigrescens View in CoL

19. Branchlets, midrib and nerves beneath and stipules puberulous; leaves with cuneate base; nervules depressed or flat above, prominent beneath; petioles 1–1.5 cm long; stipules 4–6 mm long; calyx tube c. 1 mm long, limb c. 1–1.5 mm long, more or less expanded.......................................... 109. L. sabahensis View in CoL

20. Plants spreading stiffly hirsute; leaves with subrounded and oblique base; nerves 10–12 pairs; stipules narrowly triangular or lanceolate, 4–5 mm long... 2. L. acutatus View in CoL

20. Plant otherwise.............................. 21

21. Stipules ovate or ovate-triangular or triangular with a swollen base and a membranaceous margin, 3–6 mm long 22

21. Stipules otherwise............................ 24

22. Branchlets, leaves, stipules and calyx densely spreading villose; calyx with triangular lobes; drupes villose.......................................... 58. L. hirtus View in CoL

22. Branchlets, leaf midrib and nerves beneath (on lower surface), stipules and calyx thinly puberulous or glabrous; calyx subtruncate, with 4–5 minute teeth at the apex; drupes subglabrous or glabrous................. 23

23. Branchlets, leaf midrib and nerves beneath, stipules and calyx thinly puberulous; leaves with acuminate-caudate apex; nervules obscure above, subparallel; calyx urceolate; drupes with 6 – 9 pyrenes......... 115. L. stercorarius View in CoL

23. Branchlets (except the youngest part), leaf midrib and nerves beneath and calyx quite glabrous; leaves with caudate apex; nervules distinct above, parallel; calyx shortly cupular; drupes with 4–5 pyrenes... 130. L. vulcanicus View in CoL

24. Stipules longer than 6 mm ...................... 25

24. Stipules shorter than 6 mm ..................... 29

25. Stipules broadly triangular or oblong-ovate, or oblong-lanceolate, coriaceous, glabrous or pubescent, persistent; drupes with sharp ridges....................... 26

25. Stipules lanceolate, triangular-lanceolate, linear-lanceolate or subulate, chartaceous, hirsute or strigose, often deciduous; drupes without sharp ridges................ 27

26. Stipules appressed-pubescent; leaves 12–16 by 3–4.5 cm; nerves 9–12 pairs; nervules subparallel; petioles 2– 3 mm long; calyx lobes triangular-lanceolate, 1–1.5 mm long; drupes 8–10-ridged; pyrenes 4–5.... 127. L. venosus View in CoL

26. Stipules glabrous; leaves 20 –33 by 6–11 cm; nerves 13–18 pairs; nervules reticulate; petioles 5 –10 mm long; calyx cupular with minutely teeth at the apex; drupes 6-ridged; pyrenes 6........................ 72. L. longifolius View in CoL

27. Nervules parallel; calyx lobes 4; ovary 4-locular................................... 125. L. trichophlebus View in CoL

27. Nervules reticulate to subreticulate; calyx lobes 5; ovary 5-locular................................... 28

28. Nerves 11 pairs; stipules triangular-lanceolate, 6–7 mm long, hirsute................... 90. L. palawanensis View in CoL

28. Nerves 6–8 pairs; stipules subulate, 8 mm long, appressed-pubescent....................... 91. L. papuanus View in CoL

29. Stipules 4–5 mm ............................. 30

29. Stipules shorter than 4 mm ..................... 32

30. Leaves very large, 20– 32 by 6 –10 cm; nerves 10 –12 pairs; calyx 7–8 mm long, lobes oblong, 4–5 mm long................................ 99. L. pseudolongifolius View in CoL

30. Leaves less than 20 cm long; nerves 6 –7 pairs; calyx shorter than 5 mm, with minute teeth or lobes triangular-subulate................................... 31

31. Branchlets densely appressed-pubescent; nervules parallel; calyx lobes 5 –7, triangular-subulate, as long as the tube; pyrenes 5 –7............. 114. L. solomonensis View in CoL

31. Branchlets sparsely spreading hirsute; nervules subreticulate; calyx limb minutely toothed at the apex; pyrenes 4.............................. 21. L. chlorocarpus View in CoL

32. Flowers 4-merous; calyx constricted at the base; drupes ellipsoid or ovoid, very rugose, narrowed at the base; pyrenes 1–4, deeply furrowed on the abaxial face... 33

32. Flowers 4–5-merous; calyx not constricted at base; drupes globose or subglobose, verrucose or smooth, not narrowed at the base; pyrenes 4 or more, not deeply furrowed on the abaxial face................................. 35

33. Branchlets and leaves almost glabrous; leaves chartaceous to subcoriaceous; cymes conspicuously tuberculate (short congested peduncles, up to 5 mm long); calyx with 4 trian- gular lobes; the lobes c. 1 mm long; corolla tube slender, more than 5 mm long.......... 27. L. congesticymus View in CoL

33. Branchlets and leaf nerves beneath more or less appressed puberulous, pubescent or strigose; leaves coriaceous; cymes sessile or shortly tuberculate; calyx truncate or minutely toothed or with minutely triangular lobes; corolla tube stout, less than 3 mm long................. 34

34. Branchlets thinly puberulous or sparsely strigose; leaves 8–13 by 3–4 cm; nervules 3–6 pairs; petioles 4–5 mm long; cymes sessile; calyx truncate or minutely toothed................................ 29. L. constrictus View in CoL

34. Branchlets densely appressed-pubescent; leaves 19–26 by 6– 9 cm; nerves 6 –7 pairs; petioles 5 –10; cymes shortly tuberculate; calyx with 4 minutely triangular lobes.................................... 102. L. reticulatus View in CoL

35. Calyx with a cupular limb, truncate or minutely toothed at the apex................................... 36

35. Calyx with conspicuously lobes................. 39

36. Leaves 18– 30 cm long.............. 7. L. baasianus View in CoL

36. Leaves less than 18 cm long.................... 37

37. Plant glabrous; leaves chartaceous; petioles c. 2.5 – 3.5 mm long; drupes without an enlarged cupular calyx................................... 28. L. conspicuus View in CoL

37. Branchlets and leaf venation beneath pubescent or puberulous; leaves coriaceous; petioles longer that 5 mm, drupes with an enlarged cupular calyx............ 38

38. Leaves with conspicuously reticulate nervules; nerves 10– 12 pairs; petioles 10–15 mm long; drupes 5 –6-ridged................................. 118. L. subaureus View in CoL

38. Leaves with parallel nervules; nerves 5– 9; petioles 5–10 mm long; drupes without ridges.... 128. L. verticillatus View in CoL

39. Calyx divided to the base, or calyx lobes conspicuously longer than the calyx tube...................... 40

39. Calyx not cleft to the base, calyx lobes shorter than the calyx tube or just as long as the calyx tube........ 48

40. Calyx lobes oblong, obtuse at the apex 12. L. borneensis View in CoL

40. Calyx lobes lanceolate or lanceolate-triangular, acuminate or acute at the apex.......................... 41

41. Leaves large, more than 12 cm long.... 66. L. latifolius View in CoL

41. Leaves relatively small, usually less than 12 cm long........................................... 42

42. Leaves glabrous or glabrous except appressed-hirsute or sparsely hirsute midrib and nerves and/or margin beneath; branchlets glabrous or sparsely hirsute or more or less appressed-hirsute; calyx lobes narrowly lanceolate with a long-acuminate apex or oblong; drupes glabrous or sparsely hirsute.................................. 43

42. Leaves very hairy beneath; branchlets very hairy but not appressed-hirsute; calyx lobes broadly lanceolate, usually acute at the apex; drupes hairy.................. 45

43. Leaves ovate or ovate-elliptic, usually caudate at the apex; nervules parallel; flowers 5-merous; calyx lobes narrowly lanceolate with a long acuminate apex; drupes smooth, without ridges; pyrenes 5............. 73. L. lucidus View in CoL

43. Leaves otherwise in shape; nervules reticulate; flowers 4-merous; calyx lobes lanceolate or oblong; drupes with 4 obtuse ridges; pyrenes 4..................... 44

44. Leaves oblong-lanceolate to lanceolate, 1.2–1.5 cm broad, without a marginal vein; nerves 7–9 pairs; stipules subulate-triangular, c. 1 mm long; bracts 2, subulate, 2 – 3 mm long, spreading hirsute; calyx 2 mm long, sparsely spreading hirsute; calyx lobes lanceolate, 1.2 mm long; drupes globose, smooth on the external surface...................................... 105. L. ridsdalei View in CoL

44. Leaves ovate-elliptic, 2–3.5 cm broad, with an obscure marginal vein; nerves 4–5 pairs; stipules very small, triangu- lar, c. 0.5 mm long; bracts absent; calyx c. 4.5 mm long, puberulous; calyx lobes oblong, 3 mm long; drupes subglobose, verrucose on the external surface......................................... 48. L. floresensis View in CoL

45. Leaves ovate-oblong, ovate-elliptic or ovate-lanceolate usually cuspidate at the apex; midrib glabrous above; flowers 5-merous; drupes with 5 pyrenes.......... 46

45. Leaves usually lanceolate, acute or acuminate at the apex; midrib hairy above; flowers 4-merous; drupes with 4 pyrenes.................................. 47

46. Branchlets densely tomentose; leaves ovate-oblong to oblong, tomentose beneath; nervules parallel; stipules triangular, 2 – 3 mm long; calyx not divided to the base; calyx tube c. 1 mm long; calyx lobes 1.5– 2 mm long; drupes tomentose..................... 124. L. tomentosus View in CoL

46. Branchlets densely spreading villose; leaves elliptic-lan- ceolate or ovate-lanceolate, softly villose beneath; nervules subparallel; stipules inconspicuous; calyx divided to the base; calyx lobes up to 5 mm long; drupes hirsute........................................ 35. L. curtisii View in CoL

47. Branchlets relatively slender, densely appressed-hirsute; leaves beneath and midrib above densely appressed-pubescent; bracts absent.......... 54. L. hexandrus

47. Branchlets relatively robust, densely and coarsely spread- ing hirsute; leaves beneath and midrib above densely and coarsely spreading hirsute; bracts linear, 3 – 6 mm long.............................. 25. L. chrysotrichus View in CoL

48. Branchlets and leaves glabrous and darkish when dry; leave subrounded or rounded, thickly coriaceous, 3–6 by 2–4.5 cm, rounded or subcordate at the base; venation very distinct on both surfaces; nerves 5 pairs; flowers 4-merous; calyx campanulate with 4 broadly triangular teeth; drupes with 4 pyrenes (only in Borneo)...... 108. L. rotundatus View in CoL

48. Plant otherwise.............................. 49

49. Branchlets and leaves spreading villose; leaves linear-oblong, thinly chartaceous; drupes oblate-globose, glabrous, smooth, with 7–8 obtuse ridges; pyrenes 7–8.................................. 123. L. tenuifolius View in CoL

49. Branchlets, leaves and drupes otherwise.......... 50

50. Leaves with conspicuously reticulate or subreticulate nervules...................................... 51

50. Leaves with parallel or subparallel nervules........ 64

51. Leaves lanceolate, 12–14 by 2.5–3 cm, caudate-acuminate at the apex; nerves 10–11 pairs; stipules broadly triangular, 3–4 mm, black and glabrous; calyx campanulate, 2–2.5 mm long, glabrous external surface, long yellowish hairy internal side, minutely 5– 6-toothed at the apex; drupes globose, verrucose; pyrenes 5 – 6..... 62. L. iteophyllus View in CoL

51. Plant otherwise.............................. 52

52. Branchlets and leaves beneath glabrous or thinly puberulous; leaves linear-lanceolate, with rounded and asymmetric base, 8–14 by 1.5–3 cm; petioles 1 cm long; stipules very small, triangular, c. 1 mm long; calyx limb with 5 subulate-triangular, c. 0.5 mm long teeth; drupes without ridges; pyrenes 4................. 17. L. bruneensis View in CoL

52. Plant otherwise.............................. 53

53. Branchlets glabrous or subglabrous; leaves thickly coriaceous, glabrous or subglabrous; drupes with 8 ridges 54

53. Branchlets more or less hairy; leaves chartaceous to subcoriaceous, more or less hairy on venation beneath; drupes with 4–6 ridges or without conspicuous ridges...... 55

54. Leaves lanceolate, 12–19 cm long; stipules subulate, c. 2 mm long, puberulous; calyx densely puberulous; calyx lobes ovate, c. 1 mm long; drupes puberulous........................................ 5. L. atroneurus View in CoL

54. Leaves lanceolate-oblong or oblong, 9–11 cm long; stip- ules triangular, c. 3 mm long, glabrous; calyx glabrous or sparsely puberulous; calyx lobes ovate-oblong, c. 2 mm long; drupes glabrous........... 31. L. coriaceifolius View in CoL

55. Branchlets, leaves both surfaces, stipules and calyx densely dark-brown hispid; leaves subrounded and slightly asymmetric at the base; calyx lobes 5, ovate-triangular, 1.5 by 1.5 mm, as long as the calyx tube 111. L. setulosus View in CoL

55. Plant otherwise.............................. 56

56. Leaves relatively large, longer than 15 cm, broader than 4 cm; petioles longer than 1 cm; calyx 5–7 mm long, with 4–6 triangular-ovate to orbicular reflexed lobes; drupes with 5 –6 ridges................... 20. L. chinensis View in CoL

56. Leaves less than 15 cm long and 4 cm broad; petioles less than 1 cm long; calyx less than 5 mm long; calyx lobes otherwise; drupes without ridges................. 57

57. Nerves more than 10 pairs; petioles 2 – 5 mm long... 58

57. Nerves less than 10 pairs; petioles 5 – 8 mm long.... 61

58. Branchlets and petioles appressed-pubescent or puberulous; calyx with minute teeth...... 81. L. mucronulatus View in CoL

58. Branchlets and petioles densely brownish tomentose; calyx with conspicuously lobes....................... 59

59. Leaves longer than 12 cm; nerves 14–18 pairs; petioles 5 mm long; drupes densely brownish tomentose....................................... 44. L. ferrugineus View in CoL

59. Leaves shorter than 12 cm; nerves 10–13 pairs; petioles c. 2 mm long; drupes pubescent or glabrous....... 60

60. Leaves rugose, puberulous both surfaces or only beneath, acute or caudate-cuspidate at the apex, undulate at margin; reticulate nervules obscure above; stipules subulate-triangular, densely tomentose; calyx with an obconic tube and a shortly campanulate limb..... 126. L. undulatus View in CoL

60. Leaves smooth, tomentose on midrib and nerves both surfaces, acuminate at the apex, not undulate at the margin; reticulate nervules conspicuous above; stipules broadly triangular, glabrous; calyx otherwise.. 131. L. wightianus View in CoL

61. Stipules 1.5– 3 mm long; pyrenes 6 119. L. subglobosus View in CoL

61. Stipules less than 1.5 mm long; pyrenes 4–5....... 62

62. Nerves 5 pairs; calyx campanulate; calyx lobes ovate-triangular, 1.5 –2 mm long........ 55. L. hirsutisepalus View in CoL

62. Nerves more than 6 pairs; calyx obconical or campanulate-cupular; calyx lobes triangular, 1 mm long or calyx minutely 5-toothed................................... 63

63. Branchlets densely appressed-hirsute; leaves villose beneath; midrib depressed and hirsute above; petioles densely appressed-hirsute; stipules 1 mm long, appressed-hirsute; calyx obconic, densely villose; calyx lobes triangular, 1 mm long, villose both surfaces; drupes villose. 61. L. iteoides View in CoL

63. Young branchlets puberulous, later glabrescent; leaves appressed-pubescent on midrib and nerves beneath; midrib slightly prominent and glabrous above; petioles subglabrous; stipules c. 1.5 mm long, puberulous; calyx campanulate-cupular, puberulous, minutely 5-toothed; drupes glabrous...................... 74. L. malaccensis View in CoL

64. Branchlets and leaves glabrous; leaves lanceolate, ovate-lanceolate to ovate-elliptic, cymes subsessile or shortly tuberculate; flowers extremely small; calyx c. 1 mm long .. ... ... .. ... ... ... ... ... ... ... ... ... ... ... 65

64. Plant otherwise.............................. 66

65. Leaves 8–12 by 2–3.5 cm; nerves 4–5 pairs; calyx gla- brous or sparsely puberulous; drupes oblate-globose, strongly warty; pyrenes with irregular sharp ridges.................................. 96. L. phymatodeus View in CoL

65. Leaves 15– 28 by 3.5– 5 cm; nerves 7–10 pairs; calyx densely hirsute; drupes globose, smooth; pyrenes smooth on the abaxial face.................. 79. L. minutiflorus View in CoL

66. Branchlets and leaf venation beneath densely appressed-pubescent; leaf midrib depressed above and strongly elevated beneath; calyx cupular-campanulate, 3–4 mm long; calyx lobes 5, ovate-triangular, curved inward, 1–1.5 mm long; drupes conspicuously 4–5-ridged......................................... 47. L. flavinervius View in CoL

66. Plant otherwise.............................. 67

67. Leaves relatively large, more than 5 cm broad; drupes 10–12 ridged.................... 94. L. perakensis View in CoL

67. Leaves relatively small, less than 5 cm broad; drupes with obtuse 4 ridges or smooth...................... 68

68. Leaves less than 7.5 cm long, coriaceous; flowers 4-mer- ous; drupes with 4 pyrenes..................... 69

68. Plant otherwise.............................. 70

69. Branchlets, leaf midrib and nerves beneath and petioles tomentose, calyx densely tomentose; drupes without ridges................................. 83. L. nervosus View in CoL

69. Branchlets, leaf midrib and nerves and petioles glabrous or thinly puberulous; calyx pubescent; drupes with 4 ridges................................ 82. L. myrtifolius View in CoL

70. Branchlets sparsely appressed-hirsute to glabrous; leaves coriaceous, oblong-lanceolate, 8 –13 by 2.5 – 3.5 cm, glabrous or sparsely hirsute on nerves beneath; nerves 6 – 9; nervules subparallel; stipules triangular, 3–4 mm long, coriaceous; calyx lobes 4, triangular, c. 2 mm long; drupes with 5 pyrenes................. 116. L. sterrophyllus View in CoL

70. Plant otherwise.............................. 71

71. Branchlets, leaves beneath, petioles, stipules and calyx densely brown-tomentose; leaves lanceolate or oblong-lanceolate, 9 –15 by 2 – 3.5 cm; nerves 6 –8 pairs; nervules parallel; petioles c. 1 cm long; stipules c. 2 mm long; calyx tube c. 1 mm long, the lobes triangular; drupes smooth, sparsely tomentose, with 4–5 pyrenes.. 22. L. chryseus View in CoL

71. Plant otherwise.............................. 72

72. Branchlets and leaf midrib and nerves beneath densely appressed-pubescent; leaves elliptic or oblong-elliptic, 5–10 by 1.5–4 cm, coriaceous; nerves 7–8 pairs; nervules densely anastomosing, parallel, distinct on both surfaces; stipules c. 1 mm long; flowers 4-merous; calyx limb dish-like, truncate or minutely 4-toothed; drupes with 4 pyrenes.................................. 42. L. euneurus View in CoL

72. Plant otherwise.............................. 73

73. Leaves oblong, 10–15 cm long; nerves 9 –11 pairs; petioles 1–1.5 mm long; calyx 3–4 mm long; calyx lobes triangular, 1–1.5 mm long................... 43. L. fasciculus View in CoL

73. Leaves elliptic, ovate-oblong less than 10 cm long; nerves 4–6 pairs; petioles shorter than 1 cm; calyx less than 3 mm long; calyx teeth less 1 mm long............. 74

74. Branches and leaf nerves beneath glabrous or sparsely pubescent; calyx sparsely pubescent; drupes glabrous..................................... 49. L. fordii View in CoL

74. Branches and leaves beneath densely villose or pubescent; calyx densely villous or pubescent; drupes pubescent or hirsute..................................... 75

75. Branches, leaves beneath and calyx densely spreading villose; drupes hirsute.............. 59. L. hispidulus View in CoL

75. Branches, leaves beneath and calyx more or less appressed-pubescent; drupes appressed-pubescent..................................... 26. L. clementis View in CoL