Parydra (Chaetoapnaea) nubecula Becker, 1896

Parydra (Chaetoapnaea) nubecula Becker, 1896 View in CoL

( Figs. 15 View FIGURE 15 , 89–92 View FIGURES 89–92 )

Parydra nubecula Becker 1896: 212–213 View in CoL ; type-locality: “ Schlesien “, „Patenkirchen“; syntype ♂ ♀ [ZMB, Mathis & Zatwarnicki 1995].

Primary type material examined:

♀ syntype of P. nubecula in coll. ZMB with these labels: (1) “Dohnau / 8/5 12469”; (2) “Cotypus” .

♀ syntype of P. nubecula in coll. ZMB with these labels: (1) “Kaltwasser / 12/8 24434”; (2) “Cotypus” .

♂ syntype of P. nubecula in coll. ZMB with these labels: (1) “ParTenk / 8.67”; (2) “Coll. / H. Loew ”; (3) “nubecula / Beck.”; (4) “Cotypus”; (5) “ Parydra / nubecula / Beck. / M. Krivosheina det, 1984” .

♂ syntype of P. nubecula in coll. ZMB with these labels: (1) “Dohnau / 1/5 36903”; (2) “Cotypus”; (3) “ Parydra / nubecula / Beck. / M. Krivosheina det, 1984” .

Additional material examined: 71 specimens from Germany , Finland and European Russia with new country records from Croatia and Georgia: CROATIA : 1♂ 1♀, 25.vi.2017, Ðuračica 5 km s Magić Mala [45.133°N 17.596°E] GoogleMaps ; 1♀, 29.vi.2017, floodplain Danube n Opatovac [45.264°N 19.172°E] GoogleMaps ; 1♂ 3♀♀, 26.vi.2017, small river 5.2 km sse Lacići [45.590°N 18.235°E]; GEORGIA GoogleMaps : 1♂ 1♀, 10.vii.2019, small river valley sw Manglisi [41.694°N 44.379°E] GoogleMaps .

Diagnosis: Recognised by the strongly dusted frontal vita contrasting with the subshining frontorbital plate and the characteristic wing colouration with a subapical brown band ( Fig. 15 View FIGURE 15 ). The male terminalia is uniquely characterised by the subepandrial sclerites and the shape of the phallus as described in the key and illustrated in Figs. 89–92 View FIGURES 89–92 .

Taxonomy: Mathis & Zatwarnicki (1995) incorrectly placed P. nubecula in the subgenus Parydra s. str. However, the species conforms to all characters given by Clausen & Cook (1971) for the subgenus Chaetoapnaea (new subgenus position).

Distribution in Europe: Widely distributed in Central and Eastern Europe and probably not rare even if reported only irregularly. Not recorded to date from the Iberian Peninsula, France or Britain. This may indicate an eastern distribution with the most westerly confirmed records being from Germany. Krivosheina (1989) reports P. nubecula from Northwest Russia. Records from Georgia mark the currently known southeastern distributional border. There are unexpected and isolated records from Morocco ( Pârvu et al. 2006, Popescu-Mirceni 2011) which should be checked carefully as the drawing of the wing given in one publication ( Pârvu et al. 2006: 276, 3A) does not match P. nubecula .

Biology: Typical found in wet places in forests, such as springs or around still water. Polyvoltine with the flight period spanning from March to August.