Payraudeautia profunda, Harzhauser & Landau & Guzhov, 2025

Payraudeautia profunda nov. sp.

Figs 2O View FIGURE 2 , 52A–D View FIGURE 52

Natica helicina Brocc.—Hörnes 1856: 525 View in CoL ( pars) [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Natica helicina Brocchi— Schaffer 1908 View in CoL : pl. 11, fig. 27 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Natica ( Lunatia) catena View in CoL da Costa var. helicina Brocc.— Friedberg 1923: 429 View in CoL , pl. 26, fig. 5 [ non fig. 4 = Euspira protracta View in CoL nov. sp.] [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

L [ unatia]. catena View in CoL helicina (Brocchi) View in CoL — Sieber 1958: 140 ( pars) [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Polynices ( Euspira) catena da Costa var. helicina Brocchi— Strachimirov 1960: 267 , pl. 4, figs 8–8a [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Natica ( Lunatia) catena helicina Brocchi, 1814 View in CoL — Strausz 1966: 228 ( pars), pl. 49, figs 6–8 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Lunatia catena helicina ( Brocchi, 1814) —Hinculov in Iliescu et al. 1968: 136, pl. 32, fig. 19 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Lunatia catena elatiuscula ( Sacco, 1904) —Hinculov in Iliescu et al. 1968: 136, pl. 33, fig. 1 [= non Euspira elatiuscula ( Sacco, 1904) ].

? Polinices ( Euspira) catena helicina Brocc.— Rado & Muţiu 1970: 148 , pl. 6, figs 10–11 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Lunatia catena helicina (Brocchi) — Stancu et al. 1971: 125, pl. 6, figs 8–9 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Natica ( Lunatia) catena helicina Brocchi, 1814 View in CoL — Nicorici & Sagatovici 1973: 169, pl. 25, figs 3–4 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Euspira catena helicina ( Brocchi, 1814) View in CoL — Švagrovský 1981: 141, pl. 45, figs 3, 5 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Natica ( Lunatia) helicina Brocchi, 1814 View in CoL — Krach 1981: 64, pl. 24, fig. 4 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Euspira catena helicina ( Brocchi, 1814) View in CoL — Tomašových 1998: 376, pl. 9, figs 8–10 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Type material. Holotype: NHMW 1846 View Materials /0037/0444b, SL: 16.1 mm, MD: 17.1 mm, Bad Vöslau ( Austria), Figs 52A View FIGURE 52 1 – A View FIGURE 1 4 View FIGURE 4 . Paratypes: NHMW 2024 View Materials /0236/0016, SL: 16.9 mm, MD: 17.5 mm, Bad Vöslau ( Austria), Figs 52B View FIGURE 52 1 –B View FIGURE 1 4 View FIGURE 4 . NHMW 2024 View Materials /0236/0017, SL: 18.9 mm, MD: 18.6 mm, Bad Vöslau ( Austria), Figs 52C View FIGURE 52 1 –C View FIGURE 1 4 View FIGURE 4 . NHMW 2024 View Materials /0236/0018, SL: 16.8 mm, MD: 17.8 mm, Bad Vöslau ( Austria), Figs 2O View FIGURE 2 , 52D View FIGURE 52 1 –D View FIGURE 1 4 View FIGURE 4 .

Additionalparatypes. 5spec., NHMW2024 View Materials /0236/0019, BadVöslau ( Austria) ; 1spec., NHMW1937 View Materials /0002/0308a, Bad Vöslau ( Austria) , illustrated in Schaffer (1908, pl. 11, fig. 27); 37 spec., NHMW 1846 View Materials /0037/0445, Möllersdorf ( Austria) ; 20 spec., NHMW 1862 View Materials /0001/0292, Möllersdorf ( Austria) ; 2 spec., NHMW 1862 View Materials /0001/0291, Möllersdorf ( Austria) ; 1 spec., NHMW 1865 View Materials /0001/0688, Jaroměřice ( Czech Republic) .

Type locality. Bad Vöslau ( Austria), Vienna Basin .

Type stratum. Silty clay of the Baden Formation.

Age. Middle Miocene, middle Badenian (Langhian).

Etymology. Profundus, Latin for deep, referring to the deep-water environment.

Diagnosis. Small, depressed globular shell with weakly concave, shallow subsutural ramp and wide, open umbilicus with concave umbilical wall, narrow, raised funicle, and abapical sulcus delimited by very prominent spiral ridge bent towards interior of umbilicus.

Description. Small, depressed globular shell of four teleoconch whorls; wider than high (SL/MD = 0.9); spire low, apical angle 116–131°. Protoconch low turbiniform of about three convex whorls. Suture linear, slightly impressed. Teleoconch whorls with faint subsutural cord, shallow, weakly concave subsutural ramp, delimited by slight angulation. Periphery and base convex. Last whorl inflated, attaining 90% of total height. Periphery just below mid-whorl (~47%). Growth lines distinct and weakly prosocyrt in apical view, delicate and prosocline at periphery. Aperture D-shaped, prosocline, attaining ~71% of total height. Position of adapical tip of aperture moderately low (~79%), slightly above periphery, far below shoulder. Columellar lip weakly reinforced, basal lip solid, outer lip thin. Columellar angle ~19°. Parietal callus moderately long (~45%), strongly thickened with adapical swelling and concave margin; not expanding over base. Anterior lobe moderately thick trigonal. Umbilicus moderately wide (~16%), open, deep; funicle forming narrow, raised cord deep in umbilicus with narrow adapical sulcus and wider, deeply concave abapical sulcus, delimited by very prominent spiral ridge bent towards interior of umbilicus. Umbilical callus triangular, fused with anterior lobe. Basal fasciole solid, rounded with prominent growth lines. Color pattern uniformly tan.

Discussion. This species is characterized by its depressed globose outline, weak shoulder, and the deep umbilicus with very prominent spiral ridge overhanging the umbilicus. The specimens were lumped with various Euspira species, due to their similar size and comparable color, although its umbilical features are quite distinctive. Payraudeautia ermesi Pedriali, Sosso & Dell’Angelo, 2019 differs from Payraudeautia profunda nov. sp. in the presence of a broad, depressed funicle. Payraudeautia volhynia (d’Orbigny, 1852) differs in its higher spire, depressed funicle and massive anterior lobe. All three Paratethyan species have been reported from the shallow water deposits of Varivtsi ( Ukraine), whereas Payraudeautia profunda is only known from offshore environments. Payraudeautia bituberculata Robba, Pedriali & Quaggiotto, 2016 , from the Tortonian of Tetti Borelli ( Italy), differs in its rounded knob on the parietal callus. Payraudeautia obelixi Landau, Ceulemans & Van Dingenen 2018 , from the Tortonian of France, differs in its adapical position of the aperture, and the knob-like anterior parietal lobe ( Landau et al. 2018: pl. 41, figs 1–2). Payraudeautia fasciolata ( Sacco, 1890) , from the Tortonian of Tetti Borelli and Montegibbio ( Italy), differs in its weaker parietal callus and narrower umbilical callus (see Robba et al. 2016: pl. 12, figs 2–3). Payraudeautia badensis (Glibert, 1952) differs in its high spire and the weak notch between the anterior parietal lobe and the umbilical callus.

Paleoenvironment. Occurrences in the Baden Formation of the Vienna Basin suggest middle to outer neritic settings in up to 250 m water depth ( Kranner et al. 2021).

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Friedberg 1923); Roztocze Hills: Łychów, Węglinek ( Poland) ( Krach 1981); Vienna Basin: Bad Vöslau, Möllersdorf ( Austria); Jaroměřice ( Czech Republic) ( hoc opus); Devínska Nová Ves ( Slovakia) ( Tomašových 1998); Pannonian Basin: Devecser, Szob ( Hungary) ( Strausz 1966); Dubova Region ( Romania) ( Stancu et al. 1971); Zârand Basin: Minişul des Sus ( Romania) ( Nicorici & Sagatovici 1973); Mehadia Basin : Valea Calvei, Valea Bela Reca ( Romania) (Hinculov in Iliescu et al. 1968); Dacian Basin: Islaz ( Romania) ( Rado & Muţiu 1970), Staropatitsa ( Bulgaria) ( Strachimirov 1960).