Payraudeautia volhynia (d’Orbigny, 1852), Harzhauser & Landau & Guzhov, 2025

Payraudeautia volhynia (d’Orbigny, 1852) nov. comb.

Figs 53A–F View FIGURE 53

Natica epiglottina Lam.— Dubois de Montpéreux 1831: 44 View in CoL , pl. 2, figs 34, 35 [ non Cochlis epiglottina ( Lamarck, 1804) ].

* [ Natica View in CoL ] Volhynia d’Orb.—d’Orbigny 1852: 38, nr. 576 [for Natica epiglottina sensu Dubois de Montpéreux 1831: 44 View in CoL , pl. 2, figs 34, 35].

Payraudeautia sabrinae View in CoL sp. nov. — Pedriali et al. 2019: 180 View Cited Treatment , figs 30.1−6.

Payraudeautia varovtsiana View in CoL sp. nov. — Pedriali et al. 2019: 183 View Cited Treatment , fig. 31.

non Natica Volhynica View in CoL [ sic] Orbigny– Mayer-Eymar 1895: 165 [see discussion].

Type material. Holotype, specimen illustrated in Dubois de Montpéreux (1831: pl. 2, figs 34, 35), late Badenian , Zhukivtsi ( Ukraine). We are not aware of the whereabouts of the type material, which is most probably lost.

Illustrated material. MPUM 11749, SL: 7.0 mm, MD: 6.8 mm, holotype of Payraudeautia sabrinae Pedriali, Sosso & Dell’Angelo, 2019 , Varivtsi ( Ukraine), illustrated in Pedriali et al. (2019: figs 30.1), Figs 53C View FIGURE 53 1 –C View FIGURE 1 4 View FIGURE 4 . PIN 5904/144, SL: 10.1 mm, MD: 9.2 mm, Gorodok ( Ukraine), Figs 53D View FIGURE 53 1 –D View FIGURE 1 4 View FIGURE 4 . 5904/145, SL: 7.5 mm, MD: 6.8 mm, ravine Zhabyak ( Ukraine), Figs 53E View FIGURE 53 1 –E View FIGURE 1 3 View FIGURE 3 . PIN 5904/146, Gorodok ( Ukraine), Fig. 53F View FIGURE 53 . MPUM 11754, holotype of Payraudeautia varovtsiana Pedriali, Sosso & Dell’Angelo, 2019 , SL: 5.0 mm, MD: 5.0 mm, Varivtsi ( Ukraine), illustrated in Pedriali et al. (2019: figs 31.1a−c), Figs 53A View FIGURE 53 1 –A View FIGURE 1 4 View FIGURE 4 . Paratype of Payraudeautia varovtsiana Pedriali, Sosso & Dell’Angelo, 2019 , MPUM 11755, SL: 5.2 mm, MD: 5.1 mm, Varivtsi ( Ukraine), illustrated in Pedriali et al. (2019: figs 31.2a−c), Figs 53B View FIGURE 53 1 –B View FIGURE 1 4 View FIGURE 4 .

Description. See Pedriali et al. (2019: 180).

Paratethyan synonyms. Payraudeautia sabrinae Pedriali, Sosso & Dell’Angelo, 2019 is characterized by its small, globular shell with faint shoulder angulation and massive, tongue-shaped anterior parietal lobe and prominent spiral ridge delimiting the umbilicus. These distinctive features are well depicted in the illustration drawn by Dubois de Montpéreux (1831: pl. 2, fig. 35), who had confused this small species with the French Eocene Cochlis epiglottina ( Lamarck, 1804) . The French species differs quite clearly in its semicircular umbilical callus and narrow umbilical channel (see Robba et al. 2016, pl. 1, figs 13–14). Consequently, d’Orbigny (1852) proposed Natica volhynia as name for the specimen described by Dubois de Montpéreux (1831). This name was overlooked by all subsequent authors including Pedriali et al. (2019). Therefore, we treat their P. sabrinae as subjective junior synonym of Natica volhynia d’Orbigny, 1852 . Mayer-Eymar (1895) was the first to place the Paratethyan species in Payraudeautia . Specimens identified by him as this species from the Langhian of the Loire Basin in France, however, are very unlikely to be conspecific, given the endemic character of the late Badenian fauna of the Polish-Ukrainian Foreland Basin.

Payraudeautia varovtsiana Pedriali, Sosso & Dell’Angelo, 2019 View in CoL is also treated herein as subjective junior synonym of Natica volhynia d’Orbigny, 1852 . It is characterized by its pad-like, prominent, subtrigonal anterior parietal lobe, which is separated from the semicircular umbilical callus by a notch, and by the deeply incised abapical sulcus, delimited by a sharply raised spiral ridge. Our material suggests that these features reflect the intraspecific variability of Payraudeautia volhynia .

Discussion. Pedriali et al. (2019: 180) described the protoconch as small, turbiniform, of 1.5−1.6 convex whorls. We cannot confirm this based on our material, which does not show a clear separation between protoconch and teleoconch. The earliest part of the protoconch bears delicate spirals ( Fig. 53D View FIGURE 53 5 View FIGURE 5 ).

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Late Badenian (Middle Miocene): Voronyaky Hills: Pidhirtsi ( Lviv Region, Ukraine); Ukrainian Fore-Carpathian Basin: Gorodok, Lisogirka, Sataniv, Varivtsi (Khmel’nytskyi Region), Shushkivtsi, ravine Zhabyak near Dzvynyacha, Zhukivtsi ( Ternopil Region, Ukraine) ( Dubois de Montpéreux 1831; Pedriali et al. 2019; hoc opus).

Genus Robbanacca nov. gen.

Type species. Polinices cerovaensis Harzhauser View in CoL in Harzhauser, Mandic & Schlögl, 2011, Early Miocene , Slovakia, Central Paratethys Sea .

Etymology. Combination of Robba and Nacca , in honor of Elio Robba (university of Milano, Italy) in recognition of his contributions to Neogene malacology.

Diagnosis. Small, elongate ovate shell with very low, nearly planispiral protoconch, moderately high spire, ovate last whorl; thin parietal callus with expanding, knob-like anterior parietal lobe separated from plug-like, swollen umbilical callus by deep transverse groove; umbilicus reduced to narrow chink.

Description. See description for type species.

Included species. Only the type species is known.

Discussion. Robbanacca is most similar to the Indo-West Pacific genus Polinella Marwick, 1931 [ type species Uber obstructus Marwick, 1924 , Late Miocene, New Zealand]. Polinella comprises small to medium sized, ovate shells with high last whorl and a weakly expanded parietal callus with prominent anterior parietal lobe. Its parietal callus is about the same size as the anterior lobe, filling the umbilicus to various degrees. Typically, two transverse grooves appear on the abapical part of the anterior lobe and on the umbilical callus. Species placed herein in Polinella are Uber modestus Marwick, 1924 , Uber esdailei Marwick, 1924 and Uber incertus Marwick, 1924 , all from the Cenozoic of New Zealand, but probably also Pliconacca martini Ladd, 1972 from the Early Miocene of Vita Levu on Fiji ( Ladd 1972: 8, figs 8–9) and Natica ( Lunatia) atricapilla Martin, 1884 from the Early Miocene of Indonesia ( Leloux & Wesselingh 2009: 102, pl. 202, figs 10–11). Despite the similarities with Robbanacca , the formation of the umbilical features is not homologous in the two genera. Polinella lacks a plug-like umbilicus, whereas Robbanacca lacks the transverse groove on the umbilical callus. A second group of morphologically similar species from New Zealand comprises the Early Miocene Uberella cicatrix Marwick, 1931 , the Pliocene Uberella cicatricella Marwick, 1965 and the Pleistocene to Recent Natica denticulifera Marwick, 1924 . These three species form a phylogenetic lineage of mainly bathyal species ( Beu & Maxwell 1990), which might also be close to Polinella (see Marwick 1924 and Beu & Maxwell 1990 for illustrations). Natica denticulifera Marwick, 1924 is currently listed by MolluscaBase eds. (2024) (following Spencer et al. 2209) in Uberella Finlay, 1928 [ type species Natica vitrea Hutton, 1873 , present-day, New Zealand], but the type species Uberella vitrea lacks transverse folds and has a narrow parietal callus.

Species of Polinella were placed in Pliconacca Cossmann & Martin in Martin (1914) by Ladd (1972), Majima (1989) and Robba et al. (2016). The genus Pliconacca was based on Natica ( Pliconacca) trisulcata Martin, 1914 from the Eocene of Java ( Cossmann 1925). This species has a depressed ovate outline and is characterized by a peculiar umbilical structure with a thin parietal callus and moderately thickened, expanding, semicircular umbilical callus with two long, prominent transverse folds, which do not touch each other. The most prominent, central fold coincides with the anterior parietal lobe. A third much weaker fold appears on the parietal callus and a fourth very weak fold is developed at the abapical termination of the umbilical callus. The umbilicus is reduced to a relatively wide chink (see Leloux & Wesselingh 2009: pl. 204, figs 6–15, pl. 205, figs 1–4). Therefore, it differs in shape from the elongate ovate Polinella and the formation of its umbilical features is not homologous. Similarly, the placement of the Late Miocene Natica manoharae Beets, 1942 , from Indonesia, in Pliconacca , is incorrect, because its folds are fused and heart-shaped like in Glossaulax Pilsbry, 1929 (see Beets 1942: pl. 26, figs 40–45). Glossaulax [ type species Natica reclusiana Deshayes, 1839 ; present-day, California] is distinguished from Robbanacca (as well as from Pliconacca and Polinella ) by its thick, plug-like umbilical callus composed of two completely fused lobes, separated by a narrow transverse groove. We assume that Pliconacca is restricted to the Eocene of Indonesia.

Paleoenvironment. Bathyal ( Harzhauser et al. 2011).

Distribution in Central Paratethys. Only known from the Karpatian (Early Miocene) of Cerová-Lieskové ( Slovakia) in the Vienna Basin ( Harzhauser et al. 2011).