Phaseolus coccineus

Phaseolus coccineus View in CoL L. (1753: 724) [clade B; sect. Phaseoli DC emend. Freytag ]. TYPE (LECTOTYPE): according to Westphal (1974) and Delgado-Salinas (1985) the lectotype is the specimen 899.2 in LINN, a plant cultivated in the garden of Uppsala,behaving there as an annual and of unknown origin ( McVaugh 1987).

The wild form of the scarlet runner is rather polymorphic (Delgado-Salinas 1988), namely in pubescence ( McVaugh 1987), also at molecular level ( Mina-Vargas et al. 2016); not surprisingly it has received many names ( Freytag & Debouck 2002). Some populations are clearly distinct, but possibly because of outcrossed pollination interbreeding is frequent and many intermediate forms between taxonomic entities are found. The treatment of McVaugh (1987) for “ P. coccineus and its allies” ( Piper 1926, p. 683) is followed here, excepting P. glabellus ( Fig. 6 View FIG ). While wild P. coccineus extends from Chihuahua in Mexico down to Francisco Morazán in Honduras in very diverse but open habitats at 800–3000 m, feral forms exist in El Salvador (e.g., Chalatenango: D Rodríguez 1316 (LAGU)), Costa Rica (e.g., Alajuela: A Jiménez M. 2550 (CR-118121)), Colombia (e.g., Cundinamarca: H García-Barriga 12582 (COL-30058)) and Ecuador (e.g., Pichincha: JL Luteyn 8770 (QCA)). Phaseolus multiflorus Willdenow mentioned in the list of “Vascular Plants of the Americas” ( Tropicos 2020) is a synonym. Many populations of the Central Valley of Mexico sampled by Eugène Bourgeau in 1865–1866 seem to be extinct.

Specimens: MEXICO. Chihuahua: RL Oliver 612 (DUKE-215258). Coahuila: JS Marroquín s.n. (ANSM-11433). Nuevo León: FG Meyer 2742 (BR). Tamaulipas: R Díaz 512 (RB-372771). Sinaloa: L McGill9574 (ASU-47963). Durango: S Acevedo 312 (NY). Zacatecas: JN Rose 2725 (K). San Luís Potosí: J Rzedowski 5107 (ENCB). Guanajuato: A Rubio-ChacÓn 118 (EBUM-10582). Queretaro: E Carranza 2731 (CHAPA). Hidalgo: R Hernández-Magaña 6788 (MO-3851282). Nayarit: B Bartholomew 2614 (NY). Jalisco: A Delgado-Salinas 1505 (BM). Michoacán: Ch Feddema 197 (NY). México: H Bravo H. 2006 (MA-263923). Distrito Federal: E Bourgeau 334 (G). Morelos: HS Gentry 22490 (NA). Tlaxcala: MA Martínez A. 1714 (NCU-501422). Puebla: IK Langman 2081 (PH-802749). Veracruz: E Bourgeau 3351 (P). Guerrero: DG Debouck 2359 (UC-1716201). Oaxaca: WR Anderson 13109 (DUKE-327868). Chiapas: DE Breedlove 6900 (ENCB). GUATEMALA. San Marcos: DG Debouck 1637 (M). Huehuetenango: GF Freytag 78-Guat-109 (K). Quezaltenango: DG Debouck 1675 (COL-324592). Sololá: M Nee 47366 (NY). Chimaltenango: DG Debouck 1625 (COL-324586). Sacatepéquez: WC Shannon 3683 (G). Alta Verapaz: WE Harmon 4290 (ENCB). Guatemala: A Molina R. 27546 (ENCB). El Progreso: DG Debouck 2419 (AGUAT). Santa Rosa: ET Heyde 4167 (K). Jalapa: DG Debouck 2437 (SI). HONDURAS. Ocotepeque: A Molina R. 22149 (G). Intibucá: D Dey 202 (MO-3626750). Francisco Morazán: J GÓmez 94 (BM).