Pithocerithium zelebori ( Hörnes, 1855 )

Pithocerithium zelebori ( Hörnes, 1855)

Figs 41A–E View FIGURE 41

* Cerithium Zelebori Hörn.— Hörnes 1855: 391 , pl. 41, fig. 10.

Cerithium Zelebori— Quenstedt 1884: 498 View in CoL , pl. 204, fig. 67.

? Cerithium doliolum View in CoL —Quenstedt 1884: 503, pl. 204, figs 104–105 [non Pithocerithium doliolum ( Brocchi 1814) ].? Cerithium Zelebori var. taurorara Sacc.— Sacco, 1895: 12 , pl. 1, fig. 41.

Cerithium Zelebori Hörn.— Schaffer 1912: 150 , pl. 51, figs 23–25.

C [erithium]. (Th [ericium].) zelebori Hörn.— Sieber 1958: 137 .

Cerithium (Thericium) zelebori Hoernes, 1855 — Steininger et al. 1971: 382, pl. 6, fig. 4.

non Cerithium cf. Zelebori Hörn.—Friedberg 1928: 595 , pl. 38, fig. 8 [= Pithocerithium sp. ].

non Cerithium (Thericium) aff. zelebori Hörnes, 1856 — Bałuk 1975: 147, pl. 17, figs 4–5 [= Pithocerithium longiuscatum ( Sacco, 1895) ].

non Cerithium zelebori Hörnes, 1856 — Vazzana & Cecalupo 2007: 182, pl. 5, figs a–d [= Thericium sp. ].

Type material. Lectotype designated herein: NHMW 1849 View Materials /0013/0010, SL: 36.0 mm, MD: 13.6 mm, illustrated in Hörnes (1855: pl. 41, fig. 10), Figs 41B View FIGURE 41 1 –B View FIGURE 1 2 View FIGURE 2 . Paralectotype, NHMW 2023 View Materials /0037/0001, SL: 30.6 mm, MD: 12.1 mm, Figs 41A View FIGURE 41 1 –A View FIGURE 1 2 View FIGURE 2 . Mold ( Austria), Early Miocene, Eggenburgian .

Illustrated material. NHMW 2023/0037/0002, SL: 36.0 mm, MD: 12.5 mm (largest specimen), Mold ( Austria), Figs 41C View FIGURE 41 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0037/0003, SL: 25.7 mm, MD: 11.6 mm, Mold ( Austria), Figs 41D View FIGURE 41 1 –D View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0037/0004, SL: 30. mm, MD: 10.7 mm, Mold ( Austria), Figs 41E View FIGURE 41 1 –E View FIGURE 1 2 View FIGURE 2 .

Additional material. 20 spec., NHMW 1860 View Materials /0050/0251, Mold ( Austria) , 5 spec., NHMW 2023 View Materials /0038/0001, Mold ( Austria) ; 19 spec., NHMW 2023 View Materials /0039/0001, Maria Dreieichen ( Austria) .

Synonyms. Cerithium zelebori taurorara Sacco, 1895 , from the Langhian of Italy, is highly reminiscent of P. zelebori and might be conspecific with Pithocerithium zelebori .

Revised description. Medium-sized, moderately slender conical shell of about eight teleoconch whorls, attaining ~ 30–40 mm in height; apical angle 30–35°. Protoconch and early teleoconch whorls unknown. Early spire whorls straight sided with two spiral rows of prominent, rounded tubercles. Periphery coinciding with abapical spiral cord. Interspace between spiral cords with three or more secondary spiral threads. Abapical spiral cord migrating to mid-whorl on later teleoconch whorls. Third spiral cord with weaker tubercles appears at abapical suture on penultimate whorl, mostly covered by subsequent whorl. Suture narrowly incised, undulating. Last whorl weakly convex, attaining ~45% of total height. Last whorl with three prominent spiral rows of large, rounded tubercles. Third cord of weaker, more densely spaced tubercles forms peribasal cord. Base conical, weakly constricted with three prominent spiral cords and numerous weaker spiral threads. Fasciole weak. Aperture ovate, moderately wide. Columella broadly excavated. Columellar callus forming thick, broad rim, sharply delimited from base. Anal canal weakly incised. Outer lip thin, flaring, strongly narrowing towards siphonal canal. Siphonal canal moderately long, narrow, twisted, slightly deflected to the left.

Discussion. This species is characterized by its prominent sculpture of large, rounded tubercles and its slender conical outline without marked mid-whorl angulation of the teleoconch whorls. Specimens from the Middle Miocene of the Paratethys, described by Friedberg (1928) and Bałuk (1975) as cf. and aff. zelebori , differ in their slightly ventrose outline and sculpture of smaller tubercles and might represent Pithocerithium longiuscatum ( Sacco, 1895) . Cerithium zelebori laevituberculata Sacco, 1895 , from the Burdigalian or Langhian of Italy, differs considerably in its weaker sculpture and high last whorl and is not conspecific with P. zelebori . Vazzana & Cecalupo (2007) confused specimens from the Tortonian of Cessaniti (southern Italy) with P. zelebori . These specimens have little in common with P. zelebori and lack the characteristic sculpture of tubercles. They probably represent a Thericium sp.

Paleoenvironment. Inner neritic, coastal environments in vicinity of mudflats (own data M.H.).

Distribution. Thericium zelebori is restricted to the Eggenburgian (Early Burdigalian) of the Central Paratethys Sea. The specimens from Winterlingen and Bachzimmern ( Germany), illustrated by Quenstedt (1884: pl. 204, figs 104–105) might represent additional records. The occurrence from the Langhian of Sciolze ( Italy), described by Sacco (1895), will need verification.

Central Paratethys. Eggenburgian (Early Miocene): North Alpine Foreland Basin: Eggenburg, Gauderndorf,

Maria Dreieichen, Mold, Kleinmeiseldorf ( Austria) ( Steininger et al. 1971). Eggenburgian/Ottnangian (Early

Miocene): Schwäbische Alb: Bachzimmern, Winterlingen (Swabia, Germany) (Quenstedt 1884). Proto-Mediterranean Sea.? Langhian (Middle Miocene): Po Basin: Sciolze ( Italy) ( Sacco 1895).