Pityophthorus tragardhi Spessivtsev, 1921

Pityophthorus tragardhi Spessivtsev, 1921 View in CoL

Shatilov 1987 (NE Altai)

Pityophthorus lichtensteinii (Ratzeburg, 1837) *

Material: 19: Yaylyu, 21.VII.1964, leg. AK, Pinus – 1 ex.; 20: Klyk mount, alt. 800 m, 21.VII.1964, leg. FO, Pinus – 2 ex.; 35: Chelyush cordon, alt. 1250 m, 29.VIII.1964, leg. FO, Pinus sibirica – 2 ex.

Table 1

Byctiscus betulae * FR Populus , Betula, Pyrola , Alnus , P (Linnaeus, 1758) Corylus, Tilia , Salix, Acer, Ulmus

( Voss 1930; Ter-Minassian 1950;

Koch 1992)

Byctiscus populi View in CoL *** FR Populus tremula View in CoL ; Betula View in CoL , Salix View in CoL , P (Linnaeus, 1758) Quercus View in CoL ( Voss 1930; Ter-Minassian

1950; Opanasenko & Legalov 1996)

Byctiscus rugosus * FR Populus nigra , P. balsamifera ; P (Gebler, 1829) Populus tremulae , Malus , Rubus ,

Salix ( Legalov & Legalova 2005) View in CoL

Compsapoderus View in CoL *** MD Sanguisorba View in CoL , Rosa View in CoL (Legalov & O erythropterus (Gmelin, Legalova 2005) View in CoL

1790)

Apoderus coryli View in CoL ** FR Betula View in CoL , Corylus View in CoL , Alnus View in CoL , Quercus View in CoL , Salix View in CoL P (Linnaeus, 1758) ( Ter-Minassian 1950; Opanasenko &

Legalov 1996)

Ceratapion gibbirostre * MD Carduus View in CoL ( Ehret 1990; Isaev 1994) М (Gyllenhal, 1813)

Squamapion samarense * MD Nepeta pannonica ( Ehret 1990) View in CoL М (Faust, 1891)

Protapion apricans *** * MD Trifolium ( Ehret 1990) View in CoL M (Herbst, 1797)

Protapion filirostre * MS Medicago ( Dieckmann 1977) View in CoL M (Kirby, 1808)

Protapion fulvipes *** MD Trifolium ( Arnoldi et al. 1974) View in CoL M (Geoffroy, 1785)

Catapion seniculus * MD Trifolium View in CoL ( Burakowski et al. 1992; M (Kirby, 1808) Koch 1992)

Isochnopterapion sp. * MS

Stenopterapion tenue * MS Melilotus (Dieckmann 1973) M (Kirby, 1808)

Trichapion simile ** * * FR Betula ( Opanassenko 1970; Koch М (Kirby, 1811) 1992)

Cyanapion gnarum * MF Lathyrus ( Isaev 1994) View in CoL М (Faust, 1891)

Cyanapion gyllenhali MS Vicia View in CoL (Dieckmann 1973; Arnoldi et al. М (Kirby, 1808) 1974; Burakowski et al. 1992)

Cyanapion spencii * MS Vicia ( Ehret 1990) View in CoL M (Kirby, 1808)

Eutrichapion ervi * MS Fabaceae View in CoL (Dieckmann 1973; Ehret O (Kirby, 1808) 1990; Koch 1992)

Eutrichapion viciae * * MS Vicia View in CoL , Lathyrus ( Ehret 1990) View in CoL O (Paykull, 1800)

Eutrichapion facetum * MS Vicia View in CoL , Astragalus ( Ehret 1990) View in CoL O (Gyllenhal, 1839)

Eutrichapion * MS Vicia View in CoL , Onobrychis ( Ehret 1990) View in CoL O punctigerum (Paykull,

1792)

Eutrichapion ** * MD Lathyrus ( Krivets & Korotyaev 1998) View in CoL М rhomboidale

(Desbrochers des Loges,

1871)

..continued on the next page

Table 1 Hemitrichapion sp. * - - Holotrichapion aethiops * FR Vicia ( Ehret 1990) M (Herbst, 1797) Oxystoma cerdo * ET Vicia ( Ehret 1990) M (Gerstaecker, 1854) Oxystoma opeticum *** ET Lathyrus ( Ehret 1990) M (Bach, 1854) Oxystoma subulatum * ET Lathyrus , Vicia ( Ehret 1990; Isaev O (Kirby, 1808) 1994) Notaris acridulus * WAM Glyceria ( Dieckmann 1986) M (Linnaeus, 1758) Notaris aethiops * WAM Carex ( Dieckmann 1986) M (Paykull, 1792) Notaris sp. * * * MD Cyperaceae O Thryogenes festucae * WAM Carex , Scirpus ( Dieckmann 1986) O (Herbst, 1795) Tournotaris bimaculata ** WAM Typha latifolia ( Dieckmann 1986) M (Fabricius, 1787) Dorytomus longimanus * FR Populus nigra , P. alba , P. canadensis М (Forster, 1771) ( Dieckmann 1986; Koch 1992; Legalov 2024b) Dorytomus malachovi * WA Salix ( Egorov et al. 1996) M (Korotyaev, 1976) Dorytomus suratus * FR Populus nigra ( Smreczyński 1972; М (Gyllenhal, 1835) Dieckmann 1986) Dorytomus peneckei * FR Populus tremula ( Egorov et al. 1996) М (Zumpt, 1933) Dorytomus taeniatus FR Salix ( Smreczyński 1972) М (Fabricius, 1781) Olamus (Olamus) * WA Salix ( Smreczyński 1972) M salicinus (Gyllenhal, 1827) Hylobius abietis *** * FR Pinus , rarely Abies , Picea , Larix O (Linnaeus, 1758) ( Lukjanovich & Ter-Minassian 1955; Opanassenko 1976b) Hylobius gebleri ** ** MF Bergenia crassifolia ; Rhodiola O (Boheman, 1834) ( Arnoldi et al. 1965; Egorov et al. 1996) Hylobius haroldi Faust , * FR Pinus ( Egorov et al. 1996) M 1882 Hylobius pinastri * FR Pinus , Picea , Abies , Larix (Egorov et O (Gyllenhal, 1813) al. 1996) Hylobius excavatus *** * FR Larix , other Pinaceae (Burakowski et O (Laicharting, 1781) al. 1995) Lepyrus sokolovi *** MD Probably Salix ( Krivets 1999) М (Korotyaev, 1998) Lepyrus volgensis * * * MD Salix ( Krivets 1999) М ( Faust, 1882) Pissodes castaneus * FR Pinus , rarely Abies and Larix O (DeGeer, 1775) ( Lukjanovich & Ter-Minassian 1955) Pissodes harcyniae * FR Picea , rarely Pinus , Abies and Larix O (Herbst, 1795) ( Legalov et al. 2019) ..continued on the next page

Table 1

Pissodes insignatus * FR Larix and Pinus pumila (Egorov et al. O (Boheman, 1843) 1996)

Pissodes gyllenhali * FR View in CoL Picea ( Smreczyński 1972) View in CoL M (C.R. Sahlberg, 1834)

Pissodes pini View in CoL (Linnaeus, *** * * FR Pinus View in CoL , rarely other Pinaceae (Legalov O 1758) View in CoL et al. 2019)

Pissodes piniphilus * FR Pinus , rarely other Pinaceae (Legalov O (Herbst, 1795) et al. 2019)

Pissodes validirostris * FR View in CoL Pinus ( Legalov et al. 2019) View in CoL М (Herbst, 1795)

Magdalis linearis * FR View in CoL Pinus View in CoL ( Barrios 1986; Koch 1992) М (Gyllenhal, 1827)

Magdalis phlegmatica View in CoL * * FR Pinus View in CoL , rarely Picea View in CoL and Larix View in CoL (Barrios O (Herbst, 1797) 1986; Legalov et al. 2019)

Magdalis violacea View in CoL * * FR Pinus View in CoL , Picea View in CoL ( Barrios 1986; Nikitsky O (Linnaeus, 1758) et al. 1996; Legalov et al. 2019)

Magdalis carbonaria * FR View in CoL Betula View in CoL ( Barrios 1986; Koch 1992; М (Linnaeus, 1758) Nikitsky et al. 1996)

Cryptorhynchus lapathi * FR View in CoL Salix View in CoL , Alnus View in CoL , Populus tremulae P (Linnaeus, 1758) ( Legalov et al. 2019)

Larinus impressus * MD View in CoL Asteraceae ( Ter-Minassian 1967) View in CoL O (Gebler, 1829)

Larinus iaceae View in CoL ** MD Asteraceae View in CoL ( Ter-Minassian 1967; O (Fabricius, 1775) Koch 1992; Burakowski et al. 1993)

Larinus sturnus View in CoL *** * MD Asteraceae View in CoL ( Ter-Minassian 1967; O (Schaller, 1783) Koch 1992; Burakowski et al. 1993)

Larinus turbinatus View in CoL * * MD Asteraceae View in CoL ( Ter-Minassian 1967; O (Gyllenhal, 1835) Koch 1992; Burakowski et al. 1993)

Lixus fasciculatus * MD View in CoL Artemisia ( Smreczyński 1968) М Boheman, 1835

Lixus iridis View in CoL (Olivier, *** MD Apiaceae View in CoL ( Dieckmann 1983; Koch O 1807) 1992; Burakowski et al. 1993)

Rhinocyllus conicus * MD View in CoL Cirsium View in CoL , Carduus View in CoL (Ter-Minassian O (Frölich, 1792) 1967)

Cossonus cylindricus * FR View in CoL Populus View in CoL , Salix View in CoL ( Dieckmann 1983; O (Sahlberg, 1835) Koch 1992)

Asproparthenis View in CoL * MS Amaranthaceae ( Legalov et al. 2019) View in CoL O foveocollis (Gebler,

1834)

Pseudocleonus dauricus * MD View in CoL Asteraceae ( Arnoldi et al. 1965) View in CoL O (Gebler, 1829)

Sphenophorus View in CoL * WA Cyperaceae View in CoL , Poaceae View in CoL (Arnoldi et al. P striatopunctatus View in CoL (Goeze, 1965, 1974; Smreczyński 1972;

1777) Yunakov et al. 2018)

Phloeophagus turbatus * FR View in CoL Populus View in CoL , Salix View in CoL , Betula View in CoL ( Krivets 1999; P (Schoenherr, 1845) Legalov et al. 2019)

Rhyncolus ater View in CoL ** * FR Pinaceae ( Krivets 1999) View in CoL O (Linnaeus, 1758)

Rhyncolus elongatus View in CoL ** * FR Pinaceae (Opanassenko 1976b) View in CoL O (Gyllenhal, 1827)

Rhyncolus sculpturatus * FR Pinaceae and deciduous trees (Koch P (Waltl, 1839) 1992)

..continued on the next page Table 1

Limnobaris dolorosa * WAM View in CoL Cyperaceae View in CoL , Juncaceae View in CoL ( Koch 1992; P (Goeze, 1777) Isaev 1994; Burakowski et al. 1995)

Baris przewalskyi * ST View in CoL - Zaslavskij, 1956

Dendrobaris tatjanae FR Ribes ( Egorov 1976a) View in CoL М (Egorov, 1976)

Euryommatus mariae * FR View in CoL Abies View in CoL , rarely Larix View in CoL (Egorov et al. O (Roger, 1857) 1996)

Ceutorhynchus View in CoL * FR Brassicaceae View in CoL ( Dieckmann 1972; O cochleariae View in CoL (Gyllenhal, Arnoldi et al. 1974; Koch 1992)

1813)

Ceutorhynchus View in CoL * MD Brassicaceae ( Smreczynski 1974) View in CoL O contractus (Marsham,

1802)

Ceutorhynchus typhae * MD View in CoL Brassicaceae ( Dieckmann 1972) View in CoL O (Herbst, 1795)

Ceutorhynchus View in CoL * * MD Hesperis (Korotyaev & Cholokava М pseudarator ( Korotyaev, 1989)

1989)

Ceutorhynchus matisi * MD - - Korotyaev, 1995

Glocianus punctiger * MD Taraxacum ( Koch 1992; Burakowski М (C.R. Sahlberg, 1835) et al. 1997)

Microplontus campestris * MD View in CoL Chrysanthemum leucanthemum М (Gyllenhal, 1837) ( Smreczynski 1974) View in CoL

Microplontus millefolii * MS Tanacetum View in CoL ( Dieckmann 1972; Koch М (Schultze, 1897) 1992; Burakowski et al. 1997)

Mogulones View in CoL * MD Boraginacea ( Dieckmann 1972; Koch O asperifoliarum View in CoL 1992; Burakowski et al. 1997)

(Gyllenhal, 1813)

Mogulones larvatus * MD Boraginacea ( Dieckmann 1972; O (Schultze, 1896) Smreczynski 1974)

Mogulones pallidicornis * MD Boraginacea ( Dieckmann 1972; Koch O (Gogelet & H. Brisout, 1992; Isaev 1994)

1860)

Nedyus quadrimaculatus View in CoL ** MF Urtica View in CoL ( Dieckmann 1972; Koch 1992; М (Linnaeus, 1758) Burakowski et al. 1997)

Poophagus sisymbrii * WAM View in CoL Nasturtium officinale View in CoL , Rorippa View in CoL O (Fabricius, 1777) amphibia ( Dieckmann 1972)

Ranunculiphilus View in CoL * * MF Delphinium View in CoL ( Dieckmann 1972; O inclemens (Faust, 1888) Korotyaev & Cholokava 1989 )

Svetlaniolus jurganovae * MF ??

( Korotyaev, 1980)

Thamiocolus View in CoL * MF Phlomoides ( Dieckmann 1972) View in CoL М nubeculosus (Gyllenhal,

1837)

Thamiocolus virgatus * * MS Phlomoides ( Dieckmann 1972) View in CoL М (Gyllenhal, 1837)

Trichosirocalus View in CoL * MS Asteraceae View in CoL ( Dieckmann 1972; Koch O barnevillei (Grenier, 1992; Isaev 1994; Burakowski et al.

1866) 1997)

Zacladus geranii View in CoL ** * MF Geranium View in CoL ( Dieckmann 1972; Koch М (Paykull, 1800) 1992; Isaev 1994)

..continued on the next page Table 1

Homorosoma asperum * WA Persicaria ( Korotyaev & Hong 2004) View in CoL М (Roelofs, 1875)

Rutidosoma graminosum * MF View in CoL Populus tremula View in CoL , P. alba View in CoL (Dieckmann М (Gistel, 1857) 1972)

Scleropterus verecundus View in CoL *** * FR Rubus ( Krivets 1999) View in CoL M (Faust, 1890)

Tapinotus sellatus * WA View in CoL Lysimachia View in CoL ( Dieckmann 1972; Koch М (Fabricius, 1794) 1992; Isaev 1994; Burakowski et al.

1997)

Eubrychius velutus * AQ View in CoL Myriophyllum ( Smreczynski 1974) View in CoL М (Beck, 1817)

Neophytobius hartmanni WA - - (Schultze, 1901)

Pelenomus waltoni * WAM Polygonum View in CoL ( Dieckmann 1972; Koch М (Boheman, 1843) 1992; Burakowski et al. 1997)

Rhinoncus bruchoides * MD View in CoL Polygonaceae View in CoL ( Dieckmann 1972; O (Herbst, 1784) Koch 1992; Isaev 1994; Burakowski et al. 1997)

Rhinoncus pericarpius * WAM View in CoL Polygonaceae View in CoL ( Koch 1992; Isaev O (Linnaeus, 1758) 1994; Burakowski et al. 1997)

Rhinoncus View in CoL ** WAM Polygonaceae View in CoL ( Dieckmann 1972; O perpendicularis View in CoL (Reich, Koch 1992; Isaev 1994; Burakowski

1797) et al. 1997)

Anthonomus phyllocola ** * FR Pinus , Picea (Lukjanovich & Ter- O (Herbst, 1795) Minassian 1955; Dieckmann 1988;

Koch 1992)

Anthonomus conspersus * FR View in CoL Sorbus View in CoL ( Dieckmann 1968; Koch 1992; М (Desbrochers des Loges, Burakowski et al. 1995)

1868)

Anthonomus incurvus * MF Padus , Sorbus , Cerasus (Legalov et O (Panzer, 1795) al. 2019)

Anthonomus rubi * MD View in CoL Rosaceae ( Dieckmann 1988) View in CoL O (Herbst, 1784)

Anthonomus sorbi * FR View in CoL Crataegus View in CoL , Sorbus View in CoL (Smreczynski O Germar, 1821 1972; Isaev 2007)

Furcipus rectirostris * FR Padus and other Rosaceae (Arnoldi et O (Linnaeus, 1758) al. 1974; Koch 1992)

Archarius salicivorus View in CoL ** FR Salix View in CoL ( Ter-Minassian 1956; М (Paykull, 1792) Dieckmann 1988)

Isochnus flagellum * FR Salix View in CoL ( Koch 1992; Isaev 1994) М (Erichson, 1902)

Orchestes calceatus * FR View in CoL Betula View in CoL ( Smreczyński 1976; М (Germar, 1821) Rheinheimer & Hassler 2010)

Orchestes rusci View in CoL (Herbst, * FR Betula View in CoL ( Opanassenko 1978; Koch M 1795) 1992)

Tachyerges View in CoL * FR Salix View in CoL , Betula View in CoL , Alnus View in CoL (Opanassenko P pseudostigma (Tempere, 1978; Koch 1992; Isaev 1994

1982) Burakowski et al. 1997)

Tachyerges salicis View in CoL *** FR Salix View in CoL ; Populus View in CoL ( Koch 1992; Isaev O (Linnaeus, 1758) 1994; Burakowski et al. 1997)

Tachyerges stigma View in CoL * FR Salix View in CoL , Betula View in CoL , Alnus View in CoL , Corylus View in CoL P (Germar, 1821) ( Opanassenko 1978; Koch 1992; Isaev

1994; Burakowski et al. 1997)

..continued on the next page

Table 1

Sibinia pellucens MD View in CoL Silene View in CoL ( Dieckmann 1988; Koch 1992; М (Scopoli, 1772) Isaev 1994; Burakowski et al. 1995)

Tychius meliloti * MD View in CoL Melilotus View in CoL ( Dieckmann 1988; Koch М (Stephens, 1831) 1992; Egorov et al. 1996; Burakowski et al. 1995, 1997)

Tychius picirostris * MS View in CoL Trifolium View in CoL ( Dieckmann 1988; Koch М (Fabricius, 1787) 1992; Egorov et al. 1996; Burakowski et al. 1995)

Tychius View in CoL * MS Fabaceae View in CoL ( Dieckmann 1988; Koch O quinquepunctatus View in CoL 1992; Egorov et al. 1996; Burakowski obscuripes (Korotyaev, et al. 1995)

1996)

Tychius stephensi View in CoL ** MD Trifolium View in CoL ( Arnoldi et al. 1965; Koch М (Schoenherr, 1835) 1992; Burakowski et al. 1995)

Tychius tectus (LeConte, * MS Astragalus View in CoL , Oxytropis View in CoL , Hedysarum View in CoL O 1876) ( Caldara 1990)

Cionus hortulanus View in CoL *** MD Scrophularia View in CoL , Verbascum View in CoL ( Koch 1992; O (Geoffroy, 1785) Isaev 1994; Burakowski et al. 1997)

Cionus montanus View in CoL ** MD Verbascum ( Smreczynski 1976) View in CoL М (Wingelmüller, 1914)

Cionus thapsus * MD View in CoL Verbascum View in CoL ( Koch 1992; Isaev 1994; М (Fabricius, 1792) Burakowski et al. 1997)

Cionus tuberculosus * MD View in CoL Scrophularia View in CoL ( Koch 1992; Isaev 1994; М (Scopoli, 1763) Burakowski et al. 1997)

Cleopomiarus graminis * MD View in CoL Campanula View in CoL ( Koch 1992; Isaev 1994; М (Gyllenhal, 1813) Burakowski et al. 1997)

Gymnetron melanarium * MD View in CoL Veronica View in CoL ( Koch 1992; Isaev 1994) М (Germar, 1821)

Gymnetron veronicae * MD View in CoL Veronica View in CoL ( Smreczynski 1976; Koch М (Germar, 1821) 1992; Burakowski et al. 1997)

Trichalophus biguttatus * TS View in CoL ?? (Gebler, 1832)

Trichalophus boeberi View in CoL ** * MF Ribes ( Krivets 1999) View in CoL M (Schoenherr, 1826)

Trichalophus maklini View in CoL ** ** MF Bergenia crassifolia ( Krivets 1999) View in CoL M (Faust, 1890)

Trichalophus View in CoL * MF Filipendula View in CoL , Spiraea View in CoL , Rubus View in CoL (Krivets O quadriguttatus (Gebler, 1999) View in CoL

1829)

Asiodonus opanassenkoi * MD Hedysarum coronarium (Legalov & М (Legalov, 1997) Reshetnikov 2020)

Asiodonus sajanicus * MD ?? (Korotyaev, 1998)

Donus lepidus ** * MD Hedysarum coronarium (Legalov & М (Capiomont, 1868) Reshetnikov 2020)

Metadonus * ST Atriplex ( Isaev 1994) View in CoL М distinguendus

(Boheman, 1842)

Pachypera deportata * * MD Stemmacantha , Saussurea (Krivets O (Bohaman, 1842) 1999)

Boreohypera View in CoL * ** ** MD Caryophyllaceae View in CoL ( Zaslavskij 1961; O diversipunctata Arnoldi et al. 1965 View in CoL ; Koch 1992;

(Schrank, 1798) Burakowski et al. 1995)

..continued on the next page

Table 1

Eririnomorphus rumicis * WAM Rumex ( Smreczynski 1968) View in CoL O (Linnaeus, 1758)

Hypera nigrirostris * MF View in CoL Fabaceae View in CoL ( Zaslavskij 1961; Krivets O (Fabricius, 1775) 1999)

Hypera meles View in CoL (Fabricius, *** MD Fabaceae View in CoL ( Zaslavskij 1961; Koch O 1792) 1992; Burakowski et al. 1995)

Hypera miles View in CoL (Paykull, * MS Fabaceae View in CoL ( Zaslavskij 1961; Koch O 1792) 1992; Burakowski et al. 1995)

Hypera transsylvanica * MS Medicago ( Zaslavskij 1961) View in CoL M (Petri, 1901)

Hypera misella (Faust, * * MF Fabaceae View in CoL probably Lathyrus View in CoL М 1882) ( Kippenberg 1986)

Zaslavskypera * MD Apiaceae ( Zaslavskij 1961) View in CoL O conmaculata (Herbst,

1784)

Phyllobius brevis * MS Asteraceae (Lukjanovich & Ter- O (Gyllenhal, 1834) Minassian 1955; Koch 1992;

Burakowski et al. 1993)

Phyllobius altaicus View in CoL ** * MD Geraniaceae View in CoL , Asteraceae View in CoL , P (Gebler, 1833) Polygonaceae ( Krivets 1999) View in CoL

Phyllobius oblongus * MD Deciduous View in CoL trees ( Reitter 1916; P (Linnaeus, 1758) Escherich 1923)

Phyllobius pomaceus View in CoL *** * * MF Betulaceae View in CoL , Salicaceae View in CoL , Rosaceae View in CoL , P (Gyllenhal, 1834) Grossulariacea, Geraniaceae View in CoL

( Lukjanovich & Ter-Minassian 1955;

Arnoldi et al. 1974; Koch 1992;

Burakowski et al. 1993 Krivets 1999)

Phyllobius pyri View in CoL ** MF Shrubs and deciduous trees (Koch P (Linnaeus, 1758) 1992; Burakowski et al. 1993; Egorov et al. 1996)

Phyllobius thalassinus * MD View in CoL Rosaceae View in CoL , Urticaceae View in CoL , Asteraceae View in CoL , P (Gyllenhal, 1834) Polygonaceae View in CoL , Fabaceae View in CoL , Роасеае

( Koch 1992; Isaev 1994; Krivets

1999)

Phyllobius maculicornis *** FR Deciduous trees (Lukjanovich & Ter- P (Germar, 1823) Minassian 1955; Koch 1992;

Burakowski et al. 1993; Isaev 1994)

Phyllobius virideaeris ** MF Asteraceae , Betulaceae , Salicaceae , P (Laicharting, 1781) Rosaceae , Fabaceae , Urticaceae

( Arnoldi et al. 1974; Koch 1992

Krivets 1999)

Otiorhynchus velutinus * MD View in CoL Fabaceae ( Dieckmann 1980) View in CoL O (Germar, 1823)

Otiorhynchus View in CoL *** *** *** MF Prefers Ribes nigrum View in CoL and Geranium View in CoL P grandineus View in CoL (Germar, albiftorum, and also on plants of

1823) families Pinaceae , Salicaceae ,

Betulaceae , Urticaceae , Polygonaceae ,

Ranunculaceae , Grossulariaceae ,

Rosaceae , Fabaceae , Geraniaceae ,

Onagraceae , Apiaceae , Cornaceae ,

Polemoniaceae , Asteraceae ( Krivets

1999)

Otiorhynchus oberti * * MF Poaceae , Rosaсeae, Betula (Krivets P ( Faust, 1887) 1999)

..continued on the next page

Table 1

Otiorhynchus tristis * MD View in CoL Rosaceae View in CoL , Fabaceae View in CoL , Asteraceae View in CoL P (Scopoli, 1763) ( Arnoldi et al. 1974; Koch 1992;

Burakowski et al. 1993; Isaev 1994)

Otiorhynchus ovatus ** MS Pinaceae , Betulaceae , Urticaceae , P (Linnaeus, 1758) Crassulaceae , Potygonaceae,

Apiaceae , Lamiaceae ,

Scrophulariaceae , Valerianaceae ,

Asteraceae , Rosaceae , Onagraceae ,

Fabaceae ( Krivets 1999) View in CoL

Otiorhynchus politus * FR View in CoL Asteraceae View in CoL ( Arnoldi et al. 1965; O Gyllenhal, 1834 Egorov et al. 1996; Dedyukhin 2012) Sphaeroptochus View in CoL * ST Spiraea View in CoL , Caragana ( Arnoldi 1964) View in CoL O fasciolatus (Gebler,

1829)

Centricnemus View in CoL * ST Asteraceae View in CoL , Fаbасеае, Rosaceae View in CoL , P leucogrammus View in CoL (Germar, Ranunculaceae ( Dieckmann 1980) View in CoL

1823)

Brachysomus echinatus View in CoL ** FR Salicaceae View in CoL , Rosaceae View in CoL , Fabaceae View in CoL , P (Bonsdorff, 1785) Pinaceae ( Krivets 1999) View in CoL

Eudipnus mollis View in CoL (Strom, * * * FR Pinaceae View in CoL , Betulaceae View in CoL , P 1768) Grossutariaceae, Rosaceae View in CoL , Fabaсеае,

Tiliaceae ( Krivets 1999)

Eusomus ovulum * MS Prefers View in CoL Asteraceae View in CoL (Arnoldi et al. P (Germar, 1823) 1965; Koch 1992; Burakowski et al.

1993)

Sitona ambiguus View in CoL *** MD Fabaceae View in CoL ( Koch 1992; Isaev 1994; O (Gyllenhal, 1834) Egorov et al. 1996)

Sitona cylindricollis * MS View in CoL Fabaceae View in CoL ( Arnoldi et al. 1974; O (Fahraeus, 1840) Dieckmann 1980; Koch 1992;

Burakowski et al. 1993; Isaev 1994;

Egorov et al. 1996)

Sitona hispidulus * MS View in CoL Fabaceae View in CoL ( Arnoldi et al. 1974; O (Fabricius, 1777) Dieckmann 1980; Koch 1992;

Burakowski et al. 1993; Isaev 1994)

Sitona lateralis * MS View in CoL Fabaceae View in CoL ( Arnoldi et al. 1974; O (Gyllenhal, 1834) Dieckmann 1980; Koch 1992;

Burakowski et al. 1993; Isaev 1994)

Sitona obsoletus * MD View in CoL Fabaceae View in CoL ( Arnoldi et al. 1974; O (Gmelin, 1790) Dieckmann 1980; Koch 1992;

Burakowski et al. 1993)

Sitona lineatus * MS View in CoL Fabaceae View in CoL ( Arnoldi et al. 1974; O (Linnaeus, 1758) Dieckmann 1980; Koch 1992;

Burakowski et al. 1993)

Sitona lineellus * MD View in CoL Fabaceae View in CoL ( Dieckmann 1980; Koch O (Bonsdorff, 1785) 1992; Burakowski et al. 1993; Isaev

1994)

Sitona longulus * MD View in CoL Fabaceae View in CoL ( Arnoldi et al. 1974; O (Gyllenhal, 1834) Dieckmann 1980; Koch 1992;

Burakowski et al. 1993)

Sitona sulcifrons View in CoL ** MD Trifolium View in CoL , Amoria View in CoL ( Dieckmann 1980; M (Thunberg, 1798) Krivets 1999)

..continued on the next page

Table 1

Sitona suturalis View in CoL *** MS Fabaceae View in CoL ( Dieckmann 1980; Koch O (Stephens, 1831) 1992; Burakowski et al. 1993; Isaev

1994)

Polydrusus amoenus * * MF Rosaceae , Betulaceae (Lukjanovich & P (Germar, 1823) Ter-Minassian 1955; Dieckmann

1980)

Polydrusus pilosus ** * FR Deciduous trees (Lukjanovich & Ter- P (Gredler, 1866) Minassian 1955; Dieckmann 1980;

Koch 1992)

Polydrusus corruscus FR View in CoL Salix View in CoL и Betula ( Dieckmann 1980) View in CoL P (Germar, 1823)

Polydrusus flavipes * FR View in CoL Betulaceae View in CoL , Salicaceae View in CoL , Rosaceae View in CoL , P (De Geer, 1775) Grossulariaceae ( Krivets 1999) View in CoL

Polydrusus pterygomalis * FR View in CoL Betulaceae View in CoL , Fagaceae View in CoL , Salicaceae View in CoL , P (Boheman, 1840) Rosaceae View in CoL (Lukjanovich & TerMinassian 1955; Koch 1992; Krivets

1999)

Polydrusus fulvicornis * FR View in CoL Betula View in CoL , Duschekia View in CoL , Alnus View in CoL ( Betulaceae View in CoL ) O (Fabricius, 1792) ( Dieckmann 1980; Krivets 1999)

Polydrusus tereticollis View in CoL *** * FR Deciduous trees and Pinaceae View in CoL P (DeGeer, 1775) ( Lukjanovich & Ter-Minassian 1955;

Koch 1992; Isaev 1994)

Paophilus albilaterus View in CoL ** MS Prefers Fabaceae View in CoL (Lukjanovich & Ter- P ( Faust, 1882) Minassian 1955; Arnoldi et al. 1974)

Pholicodes inauratus View in CoL *** * MD Extremely polyphagous species P (Boheman, 1833) ( Krivets 1999)

Dactylotinus globosus * *** MF Rosaceae , Rhamnaceae , P (Gebler, 1829) Grossulariaceae ( Egorov et al. 1996;

Krivets 1999)

Hylastes brunneus * FR Pinus , rarely Picea (Kryvolutskaya O Erichson, 1836 1996)

Hylastes opacus * FR Pinus , rarely Picea (Kryvolutskaya O Erichson, 1836 1996)

Hylurgops glabratus *** * ** FR Picea , rarely Pinus and Abies O (Eichhoff, 1878) (Kryvolutskaya 1996)

Hylurgops palliatus ** * * FR Pinaceae (Kryvolutskaya 1996) O (Gyllenhal, 1813)

Tomicus minor (Hartig, *** FR Pinus , rarely Picea (Kryvolutskaya O 1834) 1996)

Tomicus piniperda * * FR Pinus , Picea rarely Larix (Escherich O (Linnaeus, 1758) 1923; Kryvolutskaya 1996)

Xylechinus pilosus *** * * FR Picea , rarely Abies , Pinus , Larix O (Ratzeburg, 1837) ( Stark 1952; Kryvolutskaya 1996)

Phloeotribus spinulosus * FR Picea , rarely Larix (Kryvolutskaya O (Rey, 1883) 1996)

Carphoborus * FR Pinus , rarely Picea (Wood & Bright O cholodkovskyi 1992; Ehnström & Axelsson 2002;

(Spessivtsev, 1916) Kryvolutskaya 1996)

Dendroctonus micans *** * FR Picea , Abies , Pinus (Kryvolutskaya O (Kugelann, 1794) 1996)

Scolytus morawitzi * * FR Larix (Kryvolutskaya 1996) M (Semenov, 1902)

..continued on the next page Table 1

Scolytus ratzeburgi * * FR Betula ( Stark 1952; Kryvolutskaya М (Janson, 1856) 1996)

Polygraphus * FR Abies , rarely other Pinaceae (Stark O polygraphus (Linnaeus, 1952; Kryvolutskaya 1996)

1758)

Polygraphus subopacus * * FR Abies , rarely Pinus ( Stark 1952; O (Thomson, 1871) Kryvolutskaya 1996)

Dryocoetes alni (Georg, * FR Alnus , rarely Corylus , Fagus (Stark O 1856) 1952; Kryvolutskaya 1996)

Dryocoetes autographus * * * FR Larix , rarely Picea , Pinus , Abies O (Ratzeburg, 1837) ( Stark 1952; Kryvolutskaya 1996)

Dryocoetes baikalicus * FR Larix , rarely Pinus , Abies ( Stark 1952; O (Reitter, 1899) Kryvolutskaya 1996)

Dryocoetes ** * * FR Picea , rarely Abies, Punus , Larix O hectographus (Reitter, ( Stark 1952; Kryvolutskaya 1996)

1913)

Crypturgus cinereus * * FR Pinaceae ( Stark 1952; Kryvolutskaya O (Herbst, 1793) 1996)

Trypodendron lineatum ** * * FR Picea , Abies , rarely other Pinaceae O (Olivier, 1795) (Kryvolutskaya 1996)

Trypodendron signatum ** FR Deciduous trees ( Stark 1952) P (Fabricius, 1787)

Trypophloeus bispinulus * FR View in CoL Populus tremula ( Stark 1952) M Eggers, 1927 View in CoL

Trypophloeus granulatus * FR View in CoL Populus View in CoL , Salix View in CoL ( Stark 1952; O (Ratzeburg, 1837) Kryvolutskaya 1996)

Anisandrus dispar * FR Deciduous and coniferous trees (Stark P (Fabricius, 1792) 1952; Kryvolutskaya 1996)

Pityophthorus * FR Picea , Pinus sibirica ( Stark 1952; O micrographus (Linnaeus, Kryvolutskaya 1996)

1758)

Pityophthorus * * FR Pinus , rarely other Pinaceae (Stark O lichtensteinii (Ratzeburg, 1952; Kryvolutskaya 1996)

1837)

Pityophthorus morozovi * FR Picea ( Stark 1952; Kryvolutskaya M Spessivtsev, 1926 1996)

Pityophthorus glabratus * FR View in CoL Pinus View in CoL , rarely Larix ( Stark 1952) O Eichhoff, 1878 View in CoL

Pityophthorus tragardhi * FR View in CoL Picea View in CoL , rarely Pinus View in CoL ( Stark 1952; O Spessivtsev, 1921 Kryvolutskaya 1996)

Pityogenes bidentatus * FR Pinaceae ( Stark 1952; Kryvolutskaya O (Herbst, 1784) 1996)

Pityogenes *** * * FR Picea and other Pinaceae ( Stark 1952; O chalcographus Kryvolutskaya 1996 )

(Linnaeus, 1761)

Pityogenes conjunctus *** * FR Pinus sibirica , P. pumila , rarely Picea O (Reitter, 1887) ( Stark 1952; Kryvolutskaya 1996)

Pityogenes saalasi * FR View in CoL Picea ( Stark 1952) View in CoL М (Eggers, 1914)

Pityogenes quadridens * FR View in CoL Pinus View in CoL , rarely other Pinaceae View in CoL (Stark O (Hartig, 1834) 1952; Kryvolutskaya 1996)

Ips acuminatus *** FR Pinus , rarely other Pinaceae (Stark O (Gyllenhal, 1827) 1952; Kryvolutskaya 1996)

..continued on the next page

Table 1

Ips duplicatus (Sahlberg, **

1836)

Ips sexdentatus *** (Boerner, 1767)

Ips typographus * * (Linnaeus, 1758)

Orthotomicus laricis ** * (Fabricius, 1792)

Orthotomicus proximus *** (Eichhoff, 1867)

Orthotomicus suturalis * (Gyllenhal, 1827)

FR Picea , Pinus sibirica , rarely other O Pinaceae ( Stark 1952; Kryvolutskaya

1996)

FR Pinus , rarely other Pinaceae (Stark O 1952; Kryvolutskaya 1996)

FR Picea , Pinus , rarely other Pinaceae O (Kryvolutskaya 1996)

FR Pinus silvestris , rarely other Pinaceae O ( Stark 1952; Kryvolutskaya 1996)

FR Pinus , Picea , rarely other Pinaceae O ( Stark 1952; Kryvolutskaya 1996)

FR Pinus , Abies rarely other Pinaceae O ( Stark 1952; Kryvolutskaya 1996)

Altitude distribution: LE – low elevation, ME – middle elevation, HE – high elevation; Occurrence: 1– 3 specimens – *; 4– 6 specimens – **;> 6 specimens – ***; Biotopes: AQ – Aquatic, FR – forest, MD – meadow, MF – meadowforest, MS – meadow-steppe, WA – water-associated, WAM – water-associated-meadow, ST – steppe, TS – tundra-steppe, ET – eurytopic; Trophic groups: M – monophagous, O – oligophagous, P – polyphagous .

Discussion

The taxonomic composition of the Curculionoidea of the North-East Altai is represented by 238 species of 116 genera belonging to the following families: Nemonychidae – one species, Anthribidae – three species, Rhynchitidae – eleven species, Attelabidae – two species, Brentidae – 22 species, Curculionidae – 157 species and Scolytidae – 41 species. In total, four species ( Ceutorhynchus matisi , Homorosoma asperum , Dorytomus malachovi and Olamus salicinus ) are firstly recorded for the fauna of Altai Republic and 126 species for the fauna of the North-East Altai. The most diverse family Curculionidae is in the territory of the North-East Altai with the subfamilies Erirhininae – five species, Molytinae – 18 species, Lixinae – nine species, Dryophthorinae – one species, Cossoninae – five species, Conoderinae – 32 species, Curculioninae – 31 species and Entiminnae – 55 species.

In the studied region, 158 species of weevils are recorded only in the lowlands, 14 species only in the midlands, and the highland complex is represented by only eight species ( Table 1). In addition to the identified groups, there are species that occur in more than one zone. For example, two species ( Trichalophus maklini and Donus lepidus ) are in the midland zone, and 27 species in the midland – highland zone. In addition, 15 species are found in all altitudinal zones of the region. The species of the highland complex are represented by meadow ( Lepyrus sokolovi , Asiodonus opanassenkoi , A. sajanicus and Polydrusus fulvicornis ) and tundra forms ( Trichalophus biguttatus ) in terms of biotopic confinement. Trophically, these species are associated with Fabaceae ( Hedysarum ), Betulaceae or develop on Salix . The base of the midland complex is made up of species associated with forest biotopes (10 species), which are mono- and oligophages of conifers ( Hylobius pinastri , Pissodes harcyniae , Phloeotribus spinulosus , Carphoborus cholodkovskyi , Dendroctonus micans , Pityophthorus micrographus , Pityogenes conjunctus , P. saalasi ) and some Rosaceae ( Anthonomus conspersus и Trichalophus quadriguttatus ). The midland complex also includes species associated with grasslands. There is one species ( Notaris aethiops ) that prefers cold wet biotopes and develops on Carex along the banks of water bodies. The highland species show the greatest diversity (158 species), the meadow (43) and forest (48) species form its basis.

The steppe (26) and near-water (15) species are represented less common. The forest species are divided into inhabitants of conifer (23), deciduous (24) and conifer and deciduous polyphagous species (2). Some meadow species are associated with shrub vegetation (18), of which seven develop on Rosaceae , 3 species are monophages of Salix . The rest are restricted to herbaceous vegetation: monophages (seven species) and oligophages (27 species) on Fabaceae , Asteraceae (10 species) and Polygonaceae (six species). There are single species developing on Brassicaceae (three species), Chenopodioideae (three species), Boraginaceae (three species), Apiaceae (two species), Plantaginaceae (two species), Poaceae (two species), Cyperaceae (two species) and sedges and cereals (a species). Of particular interest are the species found throughout the profile from the lowlands to the highlands. Most of them are concentrated in the midland zone – these are bark beetles that are oligophages of coniferous trees. The other species are either polyphages, which allows them to occupy different biotopes over the entire altitude range, or monophages Betula ( Trichapion simile ), Salix ( Lepyrus volgensis ) and Carex ( Notaris sp. ).

The weevil fauna of the North-East Altai includes 225 species, with species richness decreasing with altitude ( Table 1). The greatest taxonomic diversity is found in the highland zone, due to the mosaic of forest (coniferous and deciduous forests) and open (meadow, steppe, near-water) biotopes. The midland complex, on the other hand, is not as diverse and is represented mainly by forest species – conifers and some grassland species. The poorest in terms of taxonomic diversity of weevils is the highland complex.

This altitudinal distribution of Curculionoidea in the altitudinal gradient can be explained by the fact that, although this group of beetles is widespread in all natural zones except arctic deserts, it reaches its maximum diversity in tropical regions. The decline in diversity from the lowlands to the highlands may be due to the small proportion of the most cold-resistant species in this group of beetles, as well as the lack of food plants.

In addition to the modern entomofauna of the region, two late Pleistocene entomocomplexes (Lebed and Kebezen) are known for the North-East Altai, which include 33 species from 21 genera of Curculionoidea ( Gurina et al. 2024, 2025). The majority, 24 species, are currently present in the region, while nine species ( Hemitrichapion tschernovi , Notaris altaica , Thryogenes nereis , Ceutorhynchus erysimi , Orobitis cyanea , Phyllobius contemptus , Otiorhynchus krattereri , O. af. ursus and Carphoborus teplouchovi ) are absent from the modern fauna of the North-East Altai. The absence of these species in the modern entomofauna can be explained by two main factors: firstly, the change in landscape and climatic conditions from the late Pleistocene to the present. Probably the colder and less humid conditions of the Pleistocene in the North-East Altai were suitable for the low mountain habitat of species such as Hemitrichapion tschernovi , Notaris altaica , Otiorhynchus krattereri and O. af. ursus. These species changed their ranges at the beginning of the Holocene. Secondly, the absence of other species ( Thryogenes nereis , Ceutorhynchus erysimi , Orobitis cyanea , Phyllobius contemptus and Carphoborus teplouchovi ) is probably due to their rarer occurrence.

Acknowledgements

The authors thank Dr. B.A. Korotyaev (Saint-Petersburg) for the identification some weevil species, Mr. Olaf Jaeger and Dr. Klaus-Dieter Klass ( Senckenberg Naturhistorische Sammlungen Dresden ) for providing types of Hylobius for study, and anonymous reviewers for the valuable comments that improved the manuscript. The study of A.A.G. and N.I.A. was funded by the Russian Science Foundation (grant No. 23-74-01024), and this study was partly funded by the Federal Fundamental Scientific Research Program, grant No. 1021051703269-9-1.6.12 ( R. Y.D. and A.A.L.) .

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Legalov, A.A. (2010 b) A review of the genus Teretriorhynchites (Coleoptera, Rhynchitidae) from the Russian fauna. Entomological Review, 90 (1), 59–70.

Legalov, A.A. (2015) Fossil Mesozoic and Cenozoic weevils (Coleoptera, Obrienioidea, Curculionoidea). Paleontological Journal, 49 (13), 1442–1513.

https://doi.org/10.1134/S0031030115130067

Legalov, A.A. (2017) Contribution to the knowledge of the family Nemonychidae (Coleoptera) with descriptions of new taxa. Ukrainian Journal of Ecology, 7 (2), 64–87.

https://doi.org/10.15421/2017_22

Legalov, A.A. (2018 a). Annotated key to weevils of world. Part 1. Families Nemonychidae, Anthribidae, Belidae, Ithyceridae, Rhynchitidae, Brachyceridae and Brentidae. Ukrainian Journal of Ecology, 8 (1), 780–831. https://doi.org/10.15421/2018_280

Legalov, A.A. (2018 b) Annotated key to weevils of world. Part 2. Subfamily Molytinae (Coleoptera, Curculionidae). Ukrainian Journal of Ecology, 8 (4), 340–350.

Legalov, A.A. (2018 c) Annotated key to weevils of world. Part 3. Subfamily Conoderinae (Coleoptera, Curculionidae). Ukrainian Journal of Ecology, 8 (4), 494–503.

Legalov, A.A. (2020 a) Annotated key to weevils of world. Part 4. Subfamilies Erirhininae, Dryophthorinae and Cossoninae (Curculionidae). Ukrainian Journal of Ecology, 10 (2), 319– 331.

Legalov, A.A. (2020 b) Annotated key to weevils of world. Part 5. Subfamily Entiminae (Curculionidae). Ukrainian Journal of Ecology, 10 (2), 332–346. https://doi.org/10.15421/2020_105

Legalov, A.A. (2020 c) Revised checklist of superfamily Curculionoidea (Coleoptera) from Siberia and Russian Far East. Acta Biologica Sibirica, 6, 437–549. https://doi.org/10.3897/abs.6.e59314

Legalov, A.A. (2020 d) Otiorhynchus yakovlevi Legalov, a new species in the tribe Otiorhynchini (Coleoptera: Curculionidae: Entiminae) from Altai (Western Siberia). Ecologica Montenegrina, 38, 1–7. https://doi.org/10.37828/em.2020.38.1

Legalov, A.A. (2021 a) A new species of the genus Otiorhynchus Germar, 1822 (Coleoptera: Curculionidae) from Ukok Plateau (Western Siberia, Altai). Ecologica Montenegrina, 42, 116– 120. https://doi.org/10.37828/em.2021.42.8

Legalov, A.A. (2021 b) A new species of the genus Otiorhynchus Germar, 1822 (Coleoptera: Curculionidae) from Central Altai (Western Siberia). Ecologica Montenegrina, 45, 33–36.

http://dx.doi.org/10.37828/em.2021.45.6

Legalov, A.A. (2021 c) A new species of the genus Anthonomus Germar, 1817 (Coleoptera, Curculionidae) from Altai with a list of the Russian species. Ecologica Montenegrina, 46, 67– 73. http://dx.doi.org/10.37828/em.2021.46.4

Legalov, A.A. (2021 d) Species of the genus Notaris Germar, 1817 (Coleoptera, Curculionidae) from Siberia and Russian Far East. Ecologica Montenegrina, 47, 18–27.

http://dx.doi.org/10.37828/em.2021.47.3

Legalov, A.A. (2022 a) A new species of the genus Eremochorus Zaslavskij, 1962 (Coleoptera: Curculionidae) from South-eastern Altai (Russia). Ecologica Montenegrina, 50, 28–32.

https://dx.doi.org/10.37828/em.2022.50.4

Legalov, A.A. (2022 b) A checklist of bark and ambrosia beetles (Coleoptera: Scolytidae and Platypodidae) from Siberia and the Russian Far East. Acta Biologica Sibirica, 8, 623–646.

Legalov, A.A. (2023 b) A new species of the genus Eremochorus Zaslavskij, 1962 (Coleoptera: Curculionidae) from Central Altai (Russia) with systematics remarks. Ecologica Montenegrina, 65, 76–85. https://dx.doi.org/10.37828/em.2023.65.10

Legalov, A.A. (2023 b) The genus Sphenophorus Schoenherr, 1838 (Coleoptera, Curculionidae) of Siberia, Russia. Euroasian Entomological Journal, 22 (6), 326–330.

https://doi.org/10.15298/euroasentj.22.VI.08

Legalov, A.A. (2024 a) Fossil history of bark-beetles (Coleoptera: Scolytidae) with descriptions of two new species. Historical Biology, 36 (2), 378–388.

https://doi.org/10.1080/08912963.2022.2157275

Legalov, A.A. (2024 b) Reclassification of the tribe Ellescini (Coleoptera: Curculionidae). Ecologica Montenegrina, 75, 200–241. https://doi.org/10.37828/em.2024.75.19

Legalov, A.A. (2025) New and interesting records of the supertribe Bariditae (Coleoptera: Curculionidae) from Siberia and Kazakhstan. Ecologica Montenegrina, 83, 47–59.

https://doi.org/10.37828/em.2025.83.5

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Legalov, A.A. & Reshetnikov, S.V. (2021 c) Redescription of Styphlus orthochaetinus (Reitter, 1916) from Siberia with new taxonomic data. Ecologica Montenegrina, 40, 118–124.

https://doi.org/10.37828/em.2021.40.9

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