Procloeon ( Pseudocentroptiloides ) shadini ( Kazlauskas 1964 )

Procloeon ( Pseudocentroptiloides) shadini ( Kazlauskas 1964)

( Figs 1 View FIGURES 1–8 , 18–24 View FIGURES 9–24 , 25–26 View FIGURES 25–28 , 207–209 View FIGURES 207–212 , 213–216 View FIGURES 213–216 )

Baetidae View in CoL gen. sp.: Neizvestnova-Zhadina 1931: 167 (lava).

Pseudocentroptilum? shadini Kazlauskas 1964: 170 (larva); Kluge 1995: 15 ( lectotype designation).

Centroptilum shadini : Keffermüller 1978: 99 ( ♀ imago); Keffermüller & Sowa 1984: 327 ( ♂ & ♀ imago, larva).

Pseudocentroptiloides ( Pseudocentroptiloides) shadini View in CoL : Jacob & Glazaczow 1987: 203 (larva).

Pseudocentroptiloides shadini View in CoL : Waltz & McCafferty 1989: 154 (larva, ♂ imago); Glazaczow & Klonowska-Olejnik 2009: 151 (larva, ♂ & ♀ imago, egg).

Cloeon ( Pseudocentroptiloides) nana : Kluge & Novikova 1992: 80 (subimago, ♂ & ♀ imago, egg).

Descriptions. See Keffermüller & Sowa (1984); Kluge & Novikova (1992).

Distribution. Palaearctic. Reported from France ( Chovet & Brulin 2016), Poland ( Keffermüller & Sowa 1984; Glazaczow & Klonowska-Olejnik 2009), Turkey ( Kazanci 2001), Central Russia ( Kazlauskas 1964) and Northern Urals ( Novikova 1984). Reared specimens examined by the author were collected in Lithuania and Urals; besides this, there are imagines (but not larvae) from Evenkia in Eastern Siberia ( Kluge & Novikova 1992).

Disagreement about structure of tergalii. For the first time, 15 larvae of this species were reported from river Oka by Neizvestnova-Zhadina (1931) under the name « Baetidae gen. sp.». Neizvestnova-Zhadina (1931: fig. 2) gave drawings of maxilla, labium and tergalius of the 2nd pair which was the single preserved tergalius among all examined individuals. The maxilla and the labium undoubtedly belong to P. shadini . The tergalius has the 2nd (dorsal) lamella ( Neizvestnova-Zhadina 1931: fig. 2c).

Among 8 larvae originally described by Kazlauskas (1964) as Pseudocentroptilum s hadini, only few ones had tergalius of the 1st pair only, and these tergalii had no dorsal lamella. Comparing these facts, Kazlauskas (1964) assumed that in P. s hadini tergalii of the 1st pair lack dorsal lamella, and the next tergalii have it.

Subsequent examinations of complete individuals showed that this conclusion was wrong, and actually all tergalii of this species lack dorsal lamella ( Keffermüller & Sowa 1984: 330, fig. 27).

Possibly, this disagreement was caused by the fact that Neizvestnova-Zhadina confused larvae of P. shadini with Procloeon macronyx . Kazlauskas (1964) reported larvae presumably determined by him as « Centroptilum nana Bogoescu » from the river Oka; according to his description, these larvae belong to P. macronyx . Neizvestnova-Zhadina (1931) did not report larvae of P. macronyx from the river Oka. It is not excluded, that she confused them with the larvae of P. shadini , and draw the maxilla and the labium taken from P. shadini , and the tergalius of 2nd pair taken from P. macronyx .