Pseudolinices staszici ( Friedberg, 1923 ), 2025

Pseudolinices staszici ( Friedberg, 1923) nov. comb.

Figs 31–33 View FIGURE 31 View FIGURE 32 View FIGURE 33

* Natica View in CoL ( Poliniceps [ sic]) Staszici Friedb. — Friedberg 1923: 435, pl. 26, fig. 9 [not fig. 8 = undeterminable].

Natica Staszici Friedb.— Friedberg 1938: 66 View in CoL .

Natica staszici Friedb.— Urbaniak 1974: 39 View in CoL , pl. 14, figs 2a–b.

Natica millepunctata Lam., 1822 View in CoL — Yanakevich 1980: 113, pl. 18, fig. 3 [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica helicina ( Brocchi, 1814) View in CoL — Yanakevich 1980: 113, pl. 18, fig. 4 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Natica millepunctata Lamarck, 1822 View in CoL — Yanakevich 1987: 66, pl. 4, fig. 13 [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica helicina ( Brocchi, 1814) View in CoL — Yanakevich 1987: 67, pl. 4, fig. 14 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Polinices staszici ( Friedberg, 1923) View in CoL — Pedriali et al. 2019: 185 View Cited Treatment , pl. 32.1–5.

non Polynices staszici Friedberg — Csepreghy-Meznerics 1950: 43, pl. 2, fig. 13 [= Pseudolinices empina ( De Gregorio, 1885) ].

non Natica ( Polynices) redempta staszici Friedberg, 1928 —Strauss 1966: 232, pl. 47, figs 33–35, pl. 48, figs 1–4 [= Pseudolinices empina ( De Gregorio, 1885) ].

Type material. Lectotype (designated herein), SL: 16.0 mm, MD: 16.2 mm, Tarnoruda ( Ukraine), illustrated in Friedberg (1923: pl. 26, fig. 9); we are not aware of the whereabouts of the type material.

Illustrated material. 5904/126, SL: 34.6 mm, MD: 36.8 mm, Yaseniv ( Ukraine), Middle Miocene, late Badenian, Figs 33A View FIGURE 33 1 –A View FIGURE 1 4 View FIGURE 4 . SPSU 3 /458, SL: 31.4 mm, MD: 30.6 mm, syntype of Natica eximia Eichwald, Zalistsi ( Ukraine), Middle Miocene , late Badenian, Figs 32D View FIGURE 32 1 –D View FIGURE 1 4 View FIGURE 4 . PIN 5904 View Materials /127, SL: 24.8 mm, MD: 23.2 mm, Bursuc ( Moldova), Middle Miocene, latest Badenian, Figs 32C View FIGURE 32 1 –C View FIGURE 1 4 View FIGURE 4 . PIN 5904 View Materials /128, SL: 19.8 mm, MD: 20.2 mm, Bursuc ( Moldova), Middle Miocene, latest Badenian, Figs 32D View FIGURE 32 1 –D View FIGURE 1 4 View FIGURE 4 . PIN 5904 View Materials /129, SL: 38.7 mm, MD: 37.2 mm, Ripyntsi ( Ukraine), Middle Miocene, late Badenian, Figs 32A View FIGURE 32 1 –A View FIGURE 1 4 View FIGURE 4 . PIN 5904 View Materials /130, SL: 25.4 mm, MD: 24, 7 mm, Yaseniv ( Ukraine), Middle Miocene, late Badenian, Figs 32B View FIGURE 32 1 –B View FIGURE 1 4 View FIGURE 4 . PIN 5904 View Materials /131, SL: 22.5 mm, MD: 21.2 mm, Ripyntsi ( Ukraine), Middle Miocene, late Badenian, Figs 32C View FIGURE 32 1 –C View FIGURE 1 4 View FIGURE 4 . PIN 5904 View Materials /132, SL: 20 mm, MD: 19.8 mm, Bursuc ( Moldova), Middle Miocene, latest Badenian, Figs 31A View FIGURE 31 1 –A View FIGURE 1 4 View FIGURE 4 . PIN 5904 View Materials /133, SL: 25.4 mm, MD: 25.8 mm, Bursuc ( Moldova), Middle Miocene, latest Badenian, Figs 32B View FIGURE 32 1 –B View FIGURE 1 4 View FIGURE 4 .

Revised description. Medium sized to large, solid, globose shell; about as high as wide or slightly wider ( SL /MD = 1.0). Spire moderately high turbiniform with convex whorls; apical angle 106–123°. Protoconch low turbiniform; transition into teleoconch unclear. Suture linear, adpressed. Teleoconch whorls with moderately broad subsutural ramp, delimited by rounded shoulder. Last whorl moderately expanding, slightly inflated, attaining ~90% of total height. Periphery below mid-whorl (~40%). Growth lines prominent, weakly prosocyrt in apical view, delicate, prosocline at periphery. Aperture wide, D-shaped. Position of adapical tip of aperture moderately high (~75–85%). Aperture attaining 70–78% of total height. Columellar lip strongly reinforced, basal lip slightly thickened, outer lip thin. Columellar angle ~18–33°. Opercular ridge in outer lip weak to subobsolete. Umbilicus from largely sealed by very broad, semi-circular umbilical callus with flat to slightly convex surface to half opened, restricted abaperturally by almost flat columellar callus with straight rim adapically and rounded one abapically. Rim of columellar callus gradually sunken in all cases, especially abapically, due to very broad umbilical callus separated from umbilical wall by narrow chink. Parietal callus moderately long (~44%), strongly thickened by adapical swelling, with distinctly concave margin. Anterior lobe developed to absent. Parietal callus and umbilical callus broadly fused. Basal fasciole solid, rounded. Operculum with smooth outer surface or with one distinct marginal groove along outer rim, and with flat central callus.

Discussion. Pseudolinices staszici seems to be another species endemic to the late Badenian of the Fore-Carpathian Basin. It displays considerable variability in columellar callosity. Two morphs predominate: one is characterized by its broad columellar callus filling the umbilicus completely, although some specimens of this morph have a slightly constricted columellar callus, leaving a narrow umbilical chink. A thickened and adapically expanding parietal callus is characteristic for this morph, which thus is close to Pseudolinices empina ( De Gregorio, 1885) . The second morph is characterized by a reduced columellar callus covering only about half of the umbilicus. Intermediates between these extreme morphs occur.

The apparent broad umbilical chink of a shell from the Pannonian Basin, identified as Polynices staszici by Csepreghy-Meznerics (1950) does not represent that species, but is a pathological specimen of Pseudolinices empina ( De Gregorio, 1885) .

Pseudolinices staszici differs from Pseudolinices empina in its smaller size, usually higher spire and typically wider umbilical chink (although the latter feature is variable). In addition, it lacks the characteristic subsutural spiral groove on the last teleoconch whorl, which occurs in Pseudolinices empina and P. redempta ( Michelotti, 1847) View in CoL . Pseudolinices staszici was tentatively treated as synonym of the Burdigalian Polinices proredemptus ( Sacco, 1890) by Robba et al. (2016: 196) but described as full species by Pedriali et al. (2019), with whom we agree. Pseudolinices proredemptus ( Sacco, 1890) has a wider, nearly circular umbilical callus and lacks a significant umbilical chink (see Robba et al. 2016: pl. 13, figs 4–6).

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Late Badenian (Middle Miocene): Voronyaky Hills: Golubytsya, Pidgirtsi ( Lviv Region, Ukraine); Polish-Ukrainian Fore-Carpathian Basin: Dryszczów (Nadrichne), Zgłobice ( Poland); Dibrova, Gorodok, Lisogirka, Postolivka (on river Zbruch), Ripyntsi, Smotrich (village), Stara Pisochna, Tarnoruda, Tsikova, Varivtsi, Velyka Levada, Zaichyky (Khmel’nytskyi Region), Oles’ko, Yaseniv ( Lviv Region), Shushkivtsi, Staryi Pochaiv, Zalistsi, Zboriv, Zhukivtsi ( Ternopil Region) ( Ukraine) ( Friedberg 1923; Pedriali et al. 2019; hoc opus); Moldovan Platform: Bursuc, Gîrbova ( Moldova) ( hoc opus).