Pyrophanes quadrimaculata Olivier, 1886

Pyrophanes quadrimaculata Olivier View in CoL

( Figs. 40−46 View FIGURES 40-46 )

Pyrophanes quadrimaculata Olivier View in CoL , in Baer 1886: 187; 1902: 72; 1907: 56; 1910: 48. McDermott 1966: 116. Ballantyne & Lambkin 2013: 111, Figs. 225, 226.

? Pyrophanes quadrimaculata var bimaculata Olivier, 1886: 187 . McDermott, 1966: 116.

Type. Holotype. Female. PHILIPPINES. Luzon. ( MNHN).

Other specimens examined: PHILIPPINES: Leyte: Tacoban xi.13.1944 ES Ross male (identified as Colophotia plagiata by F. McDermott ( CAS) . Mindanao : Kabasalan Zamboanga v.4.1932 HC Muzzall 3 males, female (female identified as Pyrophanes quadrimaculata var. bimaculata by F. McDermott) ( CAS) ; Dapitan Baker 2 males; ( USNM) . Samar Island 3 males Baker (two identified as Py. quadrimaculata by M. Pic) ( USNM) .

Diagnosis. One of four Pyrophanes with curved and swollen hind femora and curved hind tibiae in the male; distinguished from Py. appendiculata and Py. beccarii by the paler elytral colour which is dark marked at the apex and base only, and from Py. similissima sp. nov. which has dark markings at elytral apex only.

Description of Male. 6.6–7.9 mm long. Colour ( Figs. 40, 41, 42, 43 View FIGURES 40-46 ): pronotum, MS and MN dusky yellow, MN often pale yellow; elytra semitransparent, dusky yellow-light brown, with apical and basal dark brown areas occupying approx. 2/10 and 1/10 EL (basal darker area may appear paler than that at apex e.g. Fig. 40 View FIGURES 40-46 ); head antennae and palpi moderately dark brown; ventral aspect of thorax dingy orange, of metathorax brown in one Samar Is male; legs orange yellow with tibiae 1, 2 brownish and tarsi 1, 2 brown, tarsi 3 with apical 3 tarsomeres (Dapitan 1, Samar 2) or apical 4 (Illigan) brown; ventrites anterior to LO semitransparent appearing blotchy white because of underlying fat body; LOs may appear creamy with white margins (probably due to age); in V7 area of trough between LO halves and area posterior to LO brownish (may be an artefact); dorsal abdomen mottled whitish, T8 more orange yellow than preceding tergites. Pronotum: W/L 1.5–1.6; L/EL 0.2; L/BL 0.19; W/ GHW1.4–1.5. Head: GHW/SIW 3.7. Antennae: scape strongly expanded. FS 1 slightly longer than pedicel strongly produced laterally at apex; FS 2 longer than FS 1, FS 2–9 elongate, longer than wide and not laterally produced. Legs ( Figs. 41, 43 View FIGURES 40-46 ): MFC with up to 12 short, strong and densely packed teeth; femora 3 swollen and curved, tibiae 3 curved and expanded at their apices. Abdomen ( Figs. 44−46 View FIGURES 40-46 ): median area between LO halves in V7 appearing trough like with muscle impressions visible; PLP short and broad (L=W); incurving lobes with base slightly expanded, posterior margin rounded and outer margin slightly angulate, curving medially just posterior to the apex of the PLP and projecting medially beyond the pointed projections which are densely clothed with short hairs ( Fig. 46 View FIGURES 40-46 ); MPP very short, shallowly emarginate with posterolateral corners acute ( Fig. 46 View FIGURES 40-46 ); T8 lateral troughs with anterior area of spines and posterior area of dense hairs and short wide apically acute flanges; posterior margin angulate in midlateral areas, deep and wide median emargination ( Figs. 44 45 View FIGURES 40-46 ). Aedeagus: L/W approximately 3–4; b/a 0.8.

Remarks. Olivier (1886) based the variety bimaculata on a single female and did not attempt to distinguish between the two Pyrophanes species with similar elytral colouration viz. Py. similis and Py. quadrimaculata var. bimaculata , and given that there was no reliable association of males with this female we regard the variety bimaculata as species incertae.