Regiscolia rubida ( Gribodo, 1893 ), 2025
publication ID |
https://doi.org/10.11646/zootaxa.5700.1.1 |
publication LSID |
lsid:zoobank.org:pub:5589FAA6-35E7-4830-82A5-620339E38448 |
persistent identifier |
https://treatment.plazi.org/id/03B687D7-FFF4-FFC9-63E4-0902FA05D5F7 |
treatment provided by |
Plazi |
scientific name |
Regiscolia rubida ( Gribodo, 1893 ) |
status |
comb. nov. |
Regiscolia rubida ( Gribodo, 1893) , comb. nov.
( Figs 86–88 View FIGURE 86 View FIGURE 87 View FIGURE 88 )
Scolia ( Triscolia) haemorrhoidalis : Saussure 1880: 18; Bingham 1897: 77 (misdetermination). Triscolia haemorrhoidalis var. rubida Gribodo, 1893: 151 ( holotype, ♀, Turkestan, deposited in MSNG, Italy, not examined). Scolia View in CoL ( Scolia View in CoL , Triscolia ) flavifrons var. rubida : Dalla-Torre 1897: 160. Triscolia rubida : Micha 1927: 133. Triscolia rubida rubida : Micha 1927: 133–134. Scolia View in CoL ( Triscolia sectio Triscolia ) haemorrhoidalis : Betrem 1928: 229 (synonymised rubida Gribodo, 1893 with haemorrhoidalis
Fabricius, 1787). Scolia ( Triscolia) rubida : Betrem 1941: 118; Steinberg 1962: 119; Nagy 1965: 58; Muche 1975: 256; Grunberg 2002: 224–225
(resurrect rubida as valid species). Megascolia ( Regiscolia) rubida : Betrem & Bradley 1964: 443; Nagy 1967: 222; Bradley 1973a: 286; Osten 1992: 98; Osten
1999a: 424; Osten 2000: 546, 551, 557; Timokhanov & Kazenas 2000: 226; Gupta & Jonathan 2003: 142; Osten et al.
2003: 370; Osten 2005a: 1455, 1462; Osten 2005b: 42; Milko & Kazenas 2005: 38; Kumar & Rajmohana 2017: 5. Megascolia rubida : Nagy 1967: 222. Zazilayza rubida : Argaman 1996: 188.
Material examined. Kazakhstan. Almaty Region. 1♂, Chundzha, 2.VI.2024 ( C. Schmid-Egger leg.) [ CJBC]. Turkistan region . 1♀ 1♂, Turkistan, 12.VI.2018 (local coll. leg.) [ CJBC]. Kyzylorda region . 1♂, Kyzylorda, 1.V.2014 (local coll. leg.) [ CJBC] . Pakistan. Khyber Pakhtunkhwa. 1♂, Balakot, 12.IV.2025 ( C. Schmid-Egger leg.) [ CJBC] .
Uzbekistan. Kachkadaria province. 2♀ 2♂, Karchi, 3.VI.2015 (local coll. leg.) [ CJBC]. Surkhandary province . 2♂, Machai, 5.V.2012 ( V. Gromenko leg.) [ CJBC] . Tadjikistan. Districts under Tajikistan Central Government Jurisdiction region. 2♀ 1♂, Navobod , 25.V.2019 (local coll. leg.) [ CJBC]; 3♀, Sangvor, 4.VII.2021 (local coll. leg.) [ CJBC]. Khatlon region. 1♀, Vose, 29.V.2013 (local coll. leg.) [ CJBC] .
Diagnosis. Regiscolia rubida ( Gribodo, 1893) , comb. nov. differs from R. alecto (Smith, 1858) , comb. nov., R. azurea ( Christ, 1791) , comb. nov., R. capitata ( Fabricius, 1804) , comb. nov. and R. fulvifrons ( Saussure, 1855) , comb. nov. in females by having the apical part of scutum with a dense, uninterrupted band of punctures along the scutellum; the dorso-median area of propodeum short, approximately equal to the length of the metanotum; and the basal part of T2 with sparse punctures similar to those of the median part. In contrast, in R. azurea , R. capitata and R. fulvifrons , the apical part of scutum has sparser punctures medially or is impunctate, thus not forming a continuous band of punctures along the scutellum; the dorso-median area of propodeum is distinctly longer than the metanotum; and the basal part of T2 has finer and denser punctures forming a band, the median part being sparsely and coarsely punctate. In males, R. rubida differs by having the area around and between the posterior ocelli with very dense, tight, fine, and agglomerated punctures; and the length of the dorso-median area of propodeum nearly equal to that of the metanotum. In contrast, in R. azurea , R. alecto and R. fulvifrons , the area around and between the posterior ocelli has sparser and coarser punctures; and the length of the dorso-median area of propodeum is significantly greater than that of the metanotum. R. rubida is easily distinguished from R. capitata in males by having the frons and vertex distinctly more punctate. Whereas in R. capitata , the frons and vertex are almost entirely impunctate. In both sexes, the wings of R. bidens are bicolored, yellowish in the cells and dark elsewhere. Whereas in R. azurea , R. alecto , R. capitata , R. fulvifrons and R. splendida , the wings are entirely dark.
R. rubida differs from R. bidens ( Linnaeus, 1767) , comb. nov. and R. maculata ( Drury, 1773) , comb. nov. in females by having the pronotum with a large impunctate area close to tegula; and the basal part of scutum with dense and coarse punctures. In contrast in R. bidens and R. maculata , the pronotum and the basal part of scutum have finer, tighter and denser punctures.
R. rubida differs from R. almoraensis in females by having the pronotum with a large impunctate area close the tegula. Whereas in R. almoraensis , the pronotum is covered with coarser and sparser punctures.
R. rubida differs from R. bidens in males by having the upper margin of marginal cell significantly longer than the lower margin. Whereas in R. bidens , the upper margin of marginal cell is shorter or almost equal to the lower margin.
R. rubida differs from R. maculata ( Drury, 1773) , comb. nov. in males by having the basal third of scutum with sparse, coarse, and tight punctures; and the dorso-median area of propodeum with two protruding projections prolonged towards the rear. In contrast in R. maculata , the basal third of scutum has denser, finer, and coalescent punctures; and the dorso-median area of propodeum does not have protruding projections prolonged towards the rear.
R. rubida differs from R. almoraensis ( Gupta & Jonathan, 2003) , comb. nov. in males by having the wings bicolored, yellowish in the cells, dark on the rest; the integument almost reddish-brown; the pilosity entirely ferruginous; and the T2 and T3 marked with yellow. Whereas in R. almoraensis , the wings are almost entirely black, the yellowish area restricted to the costal margin near the marginal cell; the integument are predominantly black; the pilosity is entirely black; and the T3 is marked with orange-reddish, sometimes T4 weakly.
Redescription. Female ( Fig. 86 View FIGURE 86 ). Length 30–35 mm.
Head ( Figs 86B, C View FIGURE 86 ). Mandibles long and slender. Median mandibular tooth small and prominent, sometimes blunt. Anterior margin of clypeus rounded, without prominent lateral lobes; disk of clypeus not-domed and punctate apically and laterally; impunctate in the middle. Fissura frontalis distinct, ending in the frontal pit. Frons with numerous and dense punctures. Vertex densely punctate.
Mesosoma ( Fig. 86D View FIGURE 86 ). Pronotum with fine, tight and dense punctures, near the excavation of pronotum with a large impunctate surface. Tegula with coarse and sparse punctures basally and along scutum. Scutum with coarse and dense punctures on the basal third; with sparse and coarse punctures laterally; largely impunctate medially; with a fine, dense, and continuous band of punctures apically. Scutellum with fine and dense punctures basally, medially and apically, with sparse and coarse punctures laterally. Metanotum with fine, tight and dense punctures, with a thin impunctate central band. Dorso-lateral area propodeum with very dense, tight and fine punctures, without an impunctate area (mirror) anteriorly. Dorso-median area of propodeum with two protruding projections prolonged towards the rear; approximately equal to the length of the metanotum; with very fine, tight dense punctures. Upper plate of metapleuron impunctate on its lower half, finely and densely punctate on the rest. Anterior margin of marginal cell longer than or equal to posterior margin. Forewing with three submarginal cells. Hind tibiae spurs of similar length.
Metasoma ( Fig. 86E View FIGURE 86 ). T1 with a tubercle depressed in the middle; finely and sparsely punctate at base revealing a small impunctate area in its center; very densely and finely punctate laterally and apically. T2 with scattered and coarse punctures basally and medially; with very fine, tight and dense punctures apically. T3 almost entirely impunctate basally and medially; with dense, tight and fine punctures apically. T4 to T5 with similar punctation. S1 with dense and fine punctures laterally, mostly impunctate on the rest. S2 without distinct tubercle; with fine and dense punctures basally; with sparse and coarse punctures medially; with dense and fine punctures, in the middle with sparse punctures apically. S3 to S5 with similar punctation.
Coloration ( Fig. 86 View FIGURE 86 ). Most of the integument with reddish-brown tinge. Head marked with yellow on frons, vertex and tempora; ocellar region yellow-orange.Antennae orange-reddish. Mesosoma with scutellum often marked with yellow-orange and sometimes metanotum. Metasoma with T2 and T3 with two yellow spots more or less fused. Legs reddish. Wings bicolored, yellowish in the cells, dark on the rest. Pilosity tawny, except in the colored areas.
Male ( Fig. 87 View FIGURE 87 ). Length 24–28 mm.
Head ( Figs 87B, C View FIGURE 87 ). Inner margin of mandible with three distinct tooth (third apical tooth sometimes blunt and barely visible). Disk of clypeus with sparse punctures apically, dense and coarse punctures basally and laterally. Fissura frontalis visible reaching anterior ocelli. Frontal cross-furrow visible, its extension reaching the upper part of the ocular sinus. Frontal spatium with dense, tight and fine punctures. Anterior ocelli depressed, larger than posterior ocelli. Frons with dense, fine and tight punctures, impunctate in front of the anterior ocellus. Vertex with fine, tight and dense punctures, particularly around the posterior ocelli.
Mesosoma ( Fig. 87D View FIGURE 87 ). Pronotum with dense, tight and fine punctures over its entire surface, inter-points spaces almost coalescent. Tegula with dense and coarse punctures anteriorly and along the scutum, largely impunctate near the wing. Scutum with fine, tight and dense punctures in its basal third; with dense and fine punctures on the rest. Scutellum with fine and dense punctures basally, sparser medially and mostly impunctate apically. Metanotum with fine and dense punctures laterally, impunctate line in the middle. Dorso-lateral area of propodeum without impunctate area (mirror) anteriorly; with fine, dense and tight punctures. Dorso-median area of propodeum with two protruding projections prolonged towards the rear; with very fine, tight dense punctures. Anterior margin of the marginal cell longer than posterior margin. Forewing with three submarginal cells.
Metasoma ( Fig. 87E View FIGURE 87 ). T1 with a strong tubercle; with dense, fine and tight punctures over its entire surface. T2 with fine, tight and dense punctures, slightly sparse medially. T3-T6 similarly punctate. S1 raised in the middle, with dense punctures medially. S2 without tubercle; with dense and fine punctures basally and laterally, sparser medially and apically. S3 with sparse punctures medially, more denser laterally. S4-S6 similarly punctate.
Coloration ( Fig. 87 View FIGURE 87 ). Head reddish-brown, ocular sinus often yellowish-orange, sometimes tempora. Antennae orange-reddish except for the scape and pedicel brown. Mesosoma reddish-brown. Metasoma reddish-brown, T2- T3 marked by two yellow spots, generally fused. Wings bicolored, yellowish in the cells, dark on the rest. Pilosity tawny on head, pronotum, scutum, scutellum and last tergites; black on metanotum, propodeum and T1; yellow in the colored areas.
Genitalia. See Fig. 31H View FIGURE 31 .
Distribution. Afghanistan, India ( Himachal Pradesh, Jammu, Kashmir), Kazakhstan ( Almaty, Jambyl, Kyzylorda, Turkestan), Kyrgyzstan, Pakistan, Tajikistan, Turkmenistan, Uzbekistan ( Steinberg 1962; Timokhanov & Kazenas 2000; Osten 2000, 2005a; Osten et al. 2003; Gupta & Jonathan 2003; Milko & Kazenas 2005; Kumar & Rajmohana 2017) ( Fig. 88 View FIGURE 88 ). Steinberg (1962) cites M. rubida from Iran, with the locality given as Kuli-Tukai. This record is also mentioned by Osten et al. (2003), though with some doubt, as the locality Kuli-Tukai now appears to be untraceable. Pending further data concerning Iran, this record is not considered here. Nagy (1965) cites this species from Romania ( Tulcea), but this record appears to be doubtful and is also not considered here.
Biology. According to Timokhanov & Kazenas (2000), females of this species parasitize the larvae of Oryctes spp. ( Coleoptera , Scarabaeidae ). As specified by Milko & Kazenas (2005), in Kazakhstan R. rubida is found in habitats of Oryctes nasicornis punctipennis Motschulsky, 1860 , where dead trees and decaying stumps are present.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Regiscolia rubida ( Gribodo, 1893 )
Castagnet, Jean-Baptiste & Cabon, Flavien 2025 |
Scolia ( Triscolia ) haemorrhoidalis
Betrem, J. G. 1928: 229 |
Micha, I. 1927: 133 |
Micha, I. 1927: 133 |
Bingham, C. T. 1897: 77 |
Gribodo, G. 1893: 151 |
Saussure, H. de 1880: 18 |