Mnasicles, Zhang & Cong & Shen & Grishin, 2022

Styriodes Schaus, 1913 View in CoL , with Brownus Grishin, 2019 as its junior subjective synonym, and Remella Hemming, 1939 are subgenera of Mnasicles Godman, 1901

Genomic tree reveals that four closely related genera form a clade sister to Amblyscirtes Scudder, 1872 ( type species Hesperia vialis W. H. Edwards, 1862 ) ( Fig. 13 View Figure 13 ). Genetic diversity of this clade is comparable to that of Amblyscirtes although the wing pattern differences (from solid dark-brown to cream-striped a spotted) resulted in their classification into several small genera. We propose to treat all these species currently in four genera as congeneric, within Mnasicles Godman, 1901 ( type species Mnasicles geta Godman, 1901 ). Two genus-group names include particularly close relatives and we propose that Brownus Grishin, 2019 ( type and the only species Paratrytone browni Bell, 1959 ) is a junior subjective synonym of Styriodes Schaus, 1913 ( type species Styriodes lyco Schaus, 1913 ). We treat the two more distant ones: Styriodes and Remella Hemming, 1939 ( type species Hesperia remus Fabricius, 1798 ), as subgenera of Mnasicles , new status for both.