Rubus villicaulis, Koehler ex Weihe & Nees

33. R. villicaulis Koehler ex Weihe & Nees View in CoL , Rubi Germ. 30 (1824).

Stems robust, arching or procumbent, angled, with dense, long, patent hairs and an occasional pricklet; prickles hairy, straight, mostly deflexed. Leaflets 5, pubescent or grey-tomentose beneath; terminal leaflet orbicular-ovate to rather narrowly elliptical, acuminate, cordate at base, unevenly or coarsely serrate; petioles with strong, falcate prickles. Inflorescence large, broad, with numerous strong, straight, deflexed prickles, occasionally with a few glands and pricklets; the lower axillary branches rather numerous, long and oblique; upper branches cymose; terminal flowers subsessile. Sepals usually long-pointed, aciculate, somewhat deflexed; petals rather broad, pink or white; stamens and styles pink or white; carpels glabrous or pubescent; receptacle hairy. 2« = 28. N.W. & C. Europe. Au Br Cz?Ga Ge Hb He Ho?Ju Po Rs (W). Related species include: R. atrocaulis P. J. Mueller , Pollichia 1 6 -1 7: 163 (1859). Br Ga Ge. R. eduardii Borbäs, Österr. Bot. Zeitschr. 40: 247 (1890) ( R. villicaulis var. formanekianus Borbâs ). Ju. R. gelertii Frid. , Bot. Tidsskr. 15: 237 (1886). Br Da Ge. R. insularis F. Aresch. , Bot. Not. 1881: 158 (1881). Da Su. R. kelleri Halâcsy , Österr. Bot. Zeitschr. 40: 431 (1890). Au. R. lange! G. Jensen ex Frid. & Gelert, Bot. Tidsskr. 16: 67 (1887) ( R. atrocaulis auct. dan., non P. J. Mueller). Br D a Ge. R. magurensis E. I. Nyârâdy in Sävul., FI. Rep. Pop. Romàne 4: 908 (1956). Rm. R. ocnensis E. I. Nyârâdy in Sävul., op. cit. 907 (1956). Rm. R. seciurensis E. I. Nyârâdy in Sävul., op. cit. 906 (1956). Rm. R. septentrionalis W. C. R. Watson , Jour. Ecol. 33: 338 (1946) ( R. confinis Lindeb. , non P. J. Mueller). Br N o Su. R. subvillicaulis E. I. Nyârâdy in Sävul., FI. Rep. Pop. Romàne A. 906 (1956). Rm.