Taiwanobryum yunnanense (Enroth) Enroth, 2021

Taiwanobryum yunnanense (Enroth) Enroth , comb. nov. (Fig. 2). Basionym: Neckera yunnanensis Enroth, Hikobia 12: 3. fig. 2. 1996.

Fig. 2. Taiwanobryum

yunnanense (from G. & S.

Miehe 99-222-80). Scale

bar = 3 cm.

Fig. 3. Calyptothecium

acostatum (from G. & S.

Miehe 00-63-39). Scale bar

= 3 cm.

Specimens examined: Bhutan. Wangdue Phodrang, Punakha, Puna Tsang Chu, Chuselumba, evergreen Quercus forest with Cupressus and Pinus on E-facing slope, logging and grazing, 27°24’N 89°56’E, 2630 m a.s.l., 7.X.1999, G. & S. Miehe 99-210-54 (H); same coordinates, Quercus - Pinus forest with Cupressus emergents on W-facing slope, little disturbed, 2590 m a.s.l., 8.X.1999, G. & S. Miehe 99-213-24 (H). Wangdue Phodrang, Punakha, Upper Dabg Chu catchment, Ngoe Chu, Tsuga-Cupressus forest regenerating after fire on E-facing slope, 27°37’N 90°10’E, 2500 m a.s.l., 14.X.1999, G. & S. Miehe 99-222-80 (H, three duplicates). Tashigang, Tashiyangtse W of Risum Gompa, evergreen oak forest on SW-facing slope, grazed in winter, 27°42’N 91°27’N, 2600 m a.s.l., epiphytes on trunks, 7.VI.2000, G. & S. Miehe 00-84-40 (H). Lhuntse, Upper Kuru Chu, lower Sirsimbi Chu, E of Serchong, Quercus-Alnus-Betula secondary forest with Cupressus on SW-facing marble rock flank, 27°50’ N 91°9’ E, epiphytes, 27.I.2001, G. & S. Miehe 00-489-28 (H).

The new combination is based on a phylogenetic analysis (to be published in a forthcoming paper by the first author and colleagues) and on morphology. Taiwanobryum yunnanense resembles the more widespread T. crenulatum (Harv.) S. Olsson, Enroth & D. Quandt , but has clearly less distinct alar cells, non-decurrent leaf bases and mostly more strongly serrate leaf apices ( Enroth, 1996; Wu, 2011 a).

Taiwanobryum yunnanense was known only from Yunnan (Lu-shui, Teng-chong and Gong-shan counties) of the Gaoligongshan Range. Based on specimens at H, it grows between 1700–2700 m a.s.l., and the seven specimens from Bhutan were collected at 2500–2630 m a.s.l. In China it grows on hardwood trunks. The Bhutanese specimens mostly lack data on the substrate, but on the labels the habitats are described as hardwood or mixed hardwood-conifer forests and two of the specimens were epiphytes .

Illustrations: Enroth 1996: fig. 2; Wu 2011 (as Neckera yunnanensis ): p. 357, pl. 382, figs. 13–18.