Tanzanimyia tanzaniaensis, Freidberg & Zonstein & Friedman & T. & T., 2020

Tanzanimyia tanzaniaensis View in CoL n. sp.

( Figs 4, 10, 19, 22)

LSID: urn:lsid:zoobank.org:act:EAE96D5C-3560-499B-A711-4AF8675B2B9C .

Etymology: This species is named after Tanzania, the country where the type specimens were collected.

Diagnosis: This species is most similar to T. ugandaensis , differing from it in the following characters: It is larger (wing length 3.5–4.0 mm vs. 2.8–3.1 mm in T. ugandaensis ) and generally paler: 1 st flagellomere entirely yellow (partly blackened in T. ugandaensis ), posterior frontal seta brownish (blackish in T. ugandaensis ), brown bands on legs less conspicuous; wing pattern paler, with paler brown bullae, and with apical part, especially in cells r 2+3 and r 4+5, more reticulate.

Description: Head. Structure and coloration: 1.03–1.12× as high as long; fronto-facial angle about 100°; eye 1.09–1.25× as high as long; frons about 1.00–1.17× as wide as long; face 0.70–0.93× as high as frons length; antenna ( Fig. 4): 1 st flagellomere slightly shorter than length of face, semireniform, pointed dorsoapically; arista about 2.7× as long as 1 st flagellomere, basal 0.25 wider, yellowish, apical 0.75 blackish; pedicel about 1.5× as high as long. Chaetotaxy: Orbital setae pale brownish, anterior orbital seta about 1.8× as long as posterior orbital seta; anterior frontal seta whitish, about 0.77× as long as middle frontal seta, middle frontal seta mostly brownish, posterior frontal seta about 1.3× as long as middle frontal seta; ocellar seta white, about as long as posterior frontal seta; one preocellar fine white seta, short, about 0.25× as long as ocellar seta, slightly proclinate; postocellar setae whitish, acuminate, more or less parallel but slightly mesoclinate and proclinate, about 0.67× as long as ocellar seta; medial vertical seta about twice as long as posterior frontal seta; lateral vertical seta about 0.4× as long as medial vertical seta; two white paravertical setae 0.6–0.8× as long as postocellar seta; genal seta brown to blackish, about as long as anterior frontal seta; gena and postgena with many whitish setulae about 0.67× as long as genal seta.

Thorax. Structure: Scutum about 1.1× as long as wide; scutellum about 0.35× as long as scutum, slightly convex. Coloration and vestiture: Generally fits generic description; scutellum mostly blackish, scutellar margin at scutoscutellar suture and apex usually yellow. Chaetotaxy: Dorsocentral seta aligned closer to transverse suture than to level of postsutural supra-alar seta; acrostichal setulae in about 10 irregular rows; anterior section of anepisternum (anterior to anepisternal phragma) with about 12 fine hair-like setulae, mostly whitish and slightly lanceolate, some darker, and 5 additional but larger setulae along dorsal suture and 1 posteroventrally; posterior section of anepisternum with about 2 large and dark acuminate dorsomedial setae, and row of 5 medium-size anterior setulae mostly dark and acuminate but central setula white and lanceolate; katepisternum posterodorsally with 2 whitish and slightly lanceolate setae: 1 long and 1 short (about 0.33× as long as long seta), and about 10–15 short whitish to brownish hair-like scattered setulae and additional whitish, sometimes lanceolate, setulae: about 8 ventrally and 2–3 along anterior margin; anepimeron anterodorsally with 1 long, whitish seta, about 3× as long as 4 scattered, white, hair-like setulae; 1 post sutural supra-alar seta and 1 intra-alar seta; dorsoapical scutellar seta erect, white, about 0.33× as long as apical scutellar seta, and basal scutellar seta about twice as long as apical scutellar seta.

Legs. Coloration: Mid femur subapically with inconspicuous, pale brown, ventral band; hind femur basally with anteroventral to posteroventral brown spot and subapical dark brown band. Chaetotaxy: Forefemur with rows of long setae: setae of posteroventral row longest, brownish and acuminate; setae of posterior row dark brownish; setae of posterodorsal row mostly paler, slightly lanceolate, becoming darker and acuminate toward apex.

Wing ( Fig. 10). Length 3.8 mm. Length/width ratio about 2.4. Venation: Pterostigma about 1.2× as long as wide; vein R 1 dorsally with about 25 setulae with wide gap opposite bend of subcosta and ventrally with 3–4 setulae along posterior margin of pterostigma; vein R 4+5 sparsely setulose dorsally on basal half with 4–5 setulae all basal to crossvein R–M, and with about 5 setulae ventrally along vein. Pattern: Rays reaching costa usually generally elongate, narrow, dark with well-defined borders, except ray immediately anterior to R 4+5 blurred and petiolate; rays to posterior wing margin beyond vein M considerably wider and generally somewhat blurred; ray in cell R 4+5 split to two branches, one along apical section of vein R 4+5 and one on vein M, and with additional short apicomedial ray unconnected to adjacent rays; base of wing proximal to costal cleft and basal crossveins mostly hyaline, with more or less distinct brown ray; basal half of cell br with one ray connected to central dark area; anal lobe with three transverse brownish bands; alula centrally with one brownish transverse band. Bullae: Black large oval bulla in cell r 4+5, longer than crossvein R–M, opposite crossvein DM–Cu; brown bullae: cell r 4+5 with large elongate brown bulla proximal to crossvein DM–Cu, along vein R 4+5; cell dm with large, oval, conspicuous brown bulla along vein M, ranging from opposite to crossvein R–M and distally; cell m with small inconspicuous more or less round bulla near anteroproximal corner.

Abdomen. Coloration and chaetotaxy: Median vitta on tergites grayish to yellowish; posterior margin of tergits yellowish, sometimes narrow and inconspicuous, with whitish lanceolate setae, larger on posterior tergites; posterior margin of male tergite 5 and female tergite 6 dark brown, microtrichose, with mixed whitish lanceolate and dark acuminate setulae, and apical reclinate longer seta laterally and posteriorly with short brown setae; posterior margin of sternites usually yellowish.

Male terminalia: Not dissected. Female terminalia ( Figs 19, 22): Oviscape shiny, basal 0.33 mostly reddish brown, medially yellow, central area (about 0.5 of total length) yellow and apical 0.2 reddish brown; tergal-oviscapal measure about 3–4. Aculeus about 3.5× as long as wide ( Fig. 19), in dorsal view tip more or less pointed, with indistinct, rounded preapical shoulders; in lateral view slightly curved, without distinct projections dorsally or ventrally; two spermathecae ( Fig. 22), oval, about 1.8× as long as wide, moderately wrinkled and tuberculate.

Holotype: ♂ Tanzania: Mbeya, 35 km S Rt. A345 [9°09'S 33°31'E], 2200 m, 1.iv.1996, A. Freidberg ( SMNHTAU). The holotype is double-mounted, on a minutien pin in a plastic block, in good condition. GoogleMaps

Paratypes: 2♀, same collection data as holotype ( SMNHTAU) GoogleMaps .

Distribution: Southern Tanzania.

Biology: Unknown.