Tomosvaryella infundibula Motamedinia, Skevington & Földvari, 2023

Tomosvaryella infundibula Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:ABEB880A-DE59-4FB8-A631-A1A66F6205DA

Figs 48A–E View FIGURE 48 , 125 View FIGURE 125

Diagnosis: This species can be recognized by elongated surstyli; broadened in basal half, narrowed before apex, apex slightly broadened in dorsal view ( Fig. 48A View FIGURE 48 ); equal gonopods in height, rounded at apex; phallic guide short, broadened at base with final hook-like projection in lateral view; one of ejaculatory ducts covered by teeth in basal half ( Fig. 48D View FIGURE 48 ); ejaculatory apodeme funnel-shape ( Fig. 48C View FIGURE 48 ).

Description: MALE. Body length: 3.0 – 3.5 mm. Head. Pedicel brown with 2–3 upper bristles and 1–2 lower bristles; flagellum acuminate; brown. Face silvery pollinose. Frons, upper part close to antennae black, lower part grey pollinose; eyes touching for distance equal to 2.5 – 3 times ocellar triangle. Occiput silvery pollinose, slightly less so on upper half. Thorax. Postpronotum pale brown, with 2 – 3 short whitish bristles. Mesonotum (viewed obliquely from front) subshining black, anterior part indistinctly brown pollinose, slightly brownish also from the side. Scutellum with dorsocentral bristles weakly developed, dark, somewhat longer on frontal part. Halter, knob pale, stem dark brown. Legs. Mid coxa with 2 – 3 dark bristles. Trochanters and base of femora brown, femora dark brown, shining ventrally, femora silvery pollinose posteriorly except shiny hind femur; knees, basal 1/3 of tibiae yellow, tarsal segments pale brown, last segment dorsally darker. Ventroapical row of spines on fore femur tiny, 4–6, 8–10 short, black spines on mid femur; 6–8 spines on hind femur, no bristles posteriorly. Subapical (distal) spines on first four tibiae absent. Hind trochanter grey pollinose, without bristles on smoothly curved ventral side, ventrobasal spines absent on fore and mid femora. Hind tarsi not distinctly flattened but wider than fore and mid tarsi, first tarsomere slightly shorter than 2–5 combined; ventral bristles scrub-like. Pulvilli as long as last tarsal segment. Wing. Length: 2.8–3.1 mm. Upper margin of basal costal cell with two long distinct bristles. Fourth costal section 2–3 times as long as third costal section. Cross-vein r-m distinctly distal to middle of discal cell. Setulae on tegula absent. Abdomen. Viewed obliquely from front tergites golden-brownish pollinose, tergite 1 silvery grey pollinose with 6–8 short lateral bristles (up to 0.15 mm), sides faintly silvery on tergites 4 and 5 (the latter with larger spot, but weakly colored). Dispersed short dark bristles on tergites absent. Postabdomen in dorsal view: T6, S7 invisible; T5 twice as long as ST8. Genitalia without dissection: ST8 small, mainly shiny brown; membranous area lanceolate, shaped like a sunflower seed, vertical; surstyli relatively small, brown; hairy cerci visible without dissection in lateral view. Genitalia. Surstyli hairy, wide at base in ventral view, narrowing at 2/3 of the length and broadening at tip, both uniformly wide in lateral view and pointed at tip; epandrium short and wide; cerci symmetrically placed, small; ST8 with a narrowing membranous area towards epandrium. Genital capsule in dorsal view: epandrium and surstyli dark brown. Epandrium wider than long (MLE:MWE = 0.6). Both surstyli broadened in basal half, then strongly narrowed before apex, apex rather broadened less than basal ( Fig. 48A View FIGURE 48 ). Genital capsule in ventral view: subepandrial sclerite short and narrow, hidden behind phallic guide, without spines; gonopods symmetrical broad, apex rounded; phallus trifid, one of ejaculatory ducts covered by teeth in basal half ( Fig. 48B View FIGURE 48 ). Genital capsule in lateral view: both surstyli straight in basal half, strongly bent (right angle) towards sternite ( Figs 48D–E View FIGURE 48 ). phallic guide broadened with final thin hook-like projection, visible in lateral view ( Fig. 48D View FIGURE 48 ); ejaculatory apodeme funnel shape ( Fig. 48C View FIGURE 48 ).

FEMALE. Unknown.

Etymology: From the Latin infundibulum, funnel, in reference to the funnel-shaped ejaculatory apodeme.

Examined material: HOLOTYPE: AUSTRALIA: Western Australia: Millstream-Chichester Nat[ional] P[ark], Roebourne Road, 21°26’S, 117°9’E, 337m, sandy creek, Eucalyptus Spinifex grassland, 7–12.V.2003, C. Lambkin & D. Yeates, Malaise trap, ANIC [Bulk Sample] 2090, JSS15570 (1♂, WAM); Paratypes: AUSTRALIA: Western Australia: Millstream-Chichester National Park near Crossing Pool, 21°35’S, 117°4’E, 276m, over creek, Melaleuca riverine forest, 28.iv–5.v.2003, C. Lambkin & T. Weir, Malaise trap, JSS15965; JSS16190 (2♂, ANIC); 304m, Melaleuca riverine forest, 5–11.v.2003, C. Lambkin, D. Yeates & J. Recsei, Malaise trap, JSS15598 (1♂, ANIC); New South Wales: Crowdy Bay National Park, 40 km northeast of Taree, 31°39’S, 152°46’E, 16.iii.1981, B.J. Loudon, JSS8740 (1♂, ASCU); Queensland: JSS4188 (1♂, ZMAN) [previous syntype of T. pseudophanes ]; Carnarvon National Park, north of Ranger Station near Mount Rugged, 24°55’S, 148°0’E, near road, 12.X.2002, S. Boucher, sweeping, LEM_0016213 (1♂, LEM); Carnarvon Station, near Homestead, 24°48’S, 147°45’E, 753m, flowering herbs, 12–15.X.2014, Lambkin, Wright, J. & K. Wilson, Malaise trap, burnt 2014, 37270, CNC597136–7 (2♂, CNC); Halifax, 18°35’S, 146°17’E, IV.1920, F. Muir, CNCD5557–8 (2♂, BPBM); 1 km W Cooktown, 15°28’S 145°15’E, 12.v.1981, D.H. Colless, JSS177 (1♂, ANIC); 29 km S of Lakeland, 16°5’S, 144°47’E, 16.iv.1980, G.F. Hevel and J.A. Fortin, JSS10538 (1♂, USNM); 5 miles west Ravenshoe, 17°38’S, 145°29’E, 780m, 6.xi.1962, E.S. Ross & D.Q. Cavagnaro, JSS8475 (1♂, USNM); 6 km North of Mt. Molloy, 16°41’S, 145°20’E, 16.iv.1980, G.F. Hevel & J.A. Fortin, JSS10519 (1♂, USNM); Big Mitchell Creek Mareeba-Molloy Road, North Queensland, 16°48’S, 145°21’E, 04.v.1967, D.H. Colless, JSS135 (1♂, ANIC), Rocky Creek, 7 m [11.2 km] North Atherton, North Queensland, 17°7’S, 145°29’E, 3.V.1967, D.H. Colless, JSS8530 (1♂, ANIC); Brisbane Forest Park, top of Scrub Road, 27°25’S, 152°51’E, sclerophyll Eucalyptus forest, 04.xi.1995, Irwin & Gaimari, JSS8833, JSS8900 (2♂, INHS); Brisbane Forest Park, 27°25’S, 152°51’E, sclerophyll Eucalyptus forest, 8.xi.1995, Irwin & Gaimari, JSS8838 (1♂, INHS); Brisbane, 27°28’S, 153°01’E, 12.iii.1960, E.M. Exley, JSS8263 (1♂, QM); Brown Lake, Stradbroke Island, 27°29’S 153°25’E, 01.iii.1980, G. Daniels, JSS8918 (1♂, QM); Bunya Mountains, 26°51’S, 151°33’E, 20.v.1962, G.Monteith,JSS8260(1♂,QM);Burnett River,Bundaberg, 24°52’S, 152°21’E, 15–17.iii.1972, H. Frauca, JSS8538–9 (2♂, ANIC); Cape York, Stewart Range, 14°04’S, 143°41’E, 500m, 3.v–29.ix.1961, L.&M. Gressitt, CNCD5553 (1♂, BPBM); Carnarvon National Park, Consuelo land, 24°57’S 148°6’E, mahogany forest, 12.x.2002, M. von Tschirnhaus, JSS13951 (1♂, CNC); Carnarvon National Park, Mount Moffatt Section, 3 km SE Headquarters, 25°5’S, 148°01’E, 740m, 20.xi.1995, M.E. Irwin & S.D. Gaimari, JSS8901 (1♂, INHS); Carnarvon National Park, Mount Moffatt Section, 24°52’S, 148°01’E, 840m, next to water pool, 24.xi.1995, M.E. Irwin, Malaise trap, JSS8831, JSS8843 (2♂, INHS); Carnarvon National Park, Mount Moffatt Section, Malaise traps near Mount Moffatt, 25°04’S, 148°03’E, 19–23.i.1998, J.&A. Skevington & S.Winterton, Malaise trap, JSS2717 (1♂, QM); Carnarvon National Park, Mount Moffatt Section, Top Shelter Shed, 24°55’S, 148°03’E, 1100m, hilltop, 22.xi.1995, D.K. Yeates, JSS8310, JSS8314–6 (4♂, QM); Carnarvon Station, near Piebald Spring (CN1M1), 24°50’S, 147°45’E, 821m, Eucalyptus /Callistemon, in rocky gully, 13.xii.2010 – 15.vi.2011, C. Zwick & C. Wilson, Malaise trap, CNC576485, CNC576488 (2♂, CNC); Durham downs, Nokandra, 26°07’S, 149°04’E, 13.xi.1949, S.J. Paramonov, JSS8677 (1♂, ANIC); 29.iii–4.iv.1998, N. Power, Malaise trap, JSS7900 (1♂, QM); Noosa, 26°23’S, 153°06’E, 27.viii.1959, F.A. Perkins, JSS8254 (1♂, QM); Peawaddy Gorge, Mount Moffatt National Park, 24°56’S, 148°04’E, 1100m, 26.ii.1996, C.J. Burwell, S. Evans, JSS9745 (1♂, QM); SW Brisbane College, 27°28’S, 153°02’E, Eucalyptus , 1.ii.1961, J. L. Gressitt, CNCD5556 (1♂, BPBM); 16–23.i.1998, S. Winterton, N. Power & D. White, Malaise trap, JSS8431–3 (3♂, QM); 26.ix–3.x.1997, S. Winterton, N. Power & D. White, Malaise trap, JSS8849 (1♂, QM); Flinder’s Peak, 27°49’S, 152°48”E 10.iii.1963, G. Monteith, CNCD5569 (1♂, BPBM); Western Australia: 11 km E Marble Bar at Brockman Creek, 21°09’S, 119°52’E, 187m, damp, sandy creek bed, 2–14.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15684 (1♂, CNC); 22 km E Cobra Station, 24°13’S, 116°33’E, 381m, small dry wash, 26.iv–10.v.2003, M.E. Irwin & F.D. Parker, Malaise trap, JSS15681 (1♂, CNC); 4 miles SE by S of Minilya, 23°49’S, 114°36’E, 17.x.1970, D.H. Colless, JSS8701 (1♂, ANIC); Karijini National Park, Hamersley-Mount Bruce Road, 22°34’S, 118°18’E, 757m, dry rocky creek bed, Eucalyptus grassland, 25.iv– 14.v.2003, C. Lambkin & T. Weir, Malaise trap, JSS15968, JSS16117 (2♂, ANIC); 25.iv–14.v.2003, C. Lambkin & T. Weir, Malaise trap, JSS15986, JSS15989–90, JSS15992, JSS15994–6, JSS15998, JSS16000 (9♂, ANIC); Karijini National Park, Weano Gorge Road, 22°21’S, 118°15’E, 695m, grassy dry creek, Eucalyptus Acacia scrub, 20–25.iv.2003, C. Lambkin & T. Weir, Malaise trap, JSS16032–4 (3♂, ANIC); 25.iv–15.v.2003, C. Lambkin & T. Weir, Malaise trap, JSS16079 (1♂, ANIC); Karijini National Park: Juna Downs Road (between drying pools in rocky Turee Creek), 22°42’S, 118°25’E, 789m, 25.iv–14.v.2003, C. Lambkin & T. Weir, Malaise trap, CNCD3830, CNCD3847 (2♂, QM); Millstream-Chichester National Park, Roebourne Road, 21°26’S, 117°09’E, 337m, sandy creek, Eucalyptus Spinifex grassland, 27.iv–3.v.2003, C. Lambkin & T. Weir, Malaise trap, JSS16074, JSS16195, JSS16197, JSS16199 (4♂, ANIC).

Distribution: Australia (New South Wales, Northern Territory, Queensland, Western Australia) ( Fig. 125 View FIGURE 125 ).

Notes: Specimen JSS4188 was previously recognized as a syntype of T. pseudophanes . Based on the different shape of terminalia and external characters, we include it in T. infundibula sp. nov. The hind femur of T. infundibula sp. nov. has no bristles posteriorly while T. pseudophanes has posterior bristles. T. infundibula sp. nov. also has two long distinct bristles on the upper margin of basal costal cell. The shape of surstyli is also different in dorsal and lateral view ( Fig. 48A, D–E View FIGURE 48 ).