Tomosvaryella ledknapperensis Motamedinia, Skevington & Földvari, 2023

Tomosvaryella ledknapperensis Motamedinia, Skevington & Földvari sp. nov.

urn:lsid:zoobank.org:act:840F72E1-8137-48E6-9C99-DCB99BA8B44C

Figs 55A–E View FIGURE 55 , 128 View FIGURE 128

Diagnosis: This species can be recognized by the hind trochanter having a keel covered with small teeth along its edge; elongated surstyli, widened at the base and bent toward each other at the apex in dorsal view ( Fig. 55A View FIGURE 55 ); both surstyli bent towards sternite in apical third in lateral view (Fig. D–E); subepandrial sclerite with fin-shaped projection in its center, both gonopods elongated and pointed at the apex, hypandrial apodeme distinct in ventral view ( Fig. 55C View FIGURE 55 ).

Description: MALE: Body length: 2.5 mm. Head. Scape, pedicel dark brown, flagellum light yellow, pollinose. Pedicel with three short upper bristles. Flagellum tapering. Face silvery pollinose. Frons, upper part shining black; lower part distinctly silver pollinose; eyes touching for distance equal to 2 times the length of ocellar triangle. Occiput silvery pollinose, less so on upper 1/3 with scattered short dark bristles. Thorax. Postpronotum pale, with 2–3 long pale bristles. Mesonotum (viewed obliquely from front) silvery pollinose, anterior part more grey, grayish also from the side. Scutellum silvery brown pollinose with 2–4 short bristles along distal edge. Halter, knob and tip light brown. Legs. Mid coxa with 3–4 different-size dark bristles. Trochanters light brown, femora brown, silvery pollinose posteriorly except shiny hind femur; knees and basal 1/6 of tibiae light yellow (tibiae otherwise light brown), tarsal segments bright yellow completely. Hind trochanter with a keel covering small teeth along its edge in a row; ventrobasal spines present on fore and mid femora. Hind femur without ventroapical spines. Subapical (distal) spines on first four tibiae present (short). Hind tarsomeres not flattened, hind metatarsus almost as long as 2–4 combined, dorsally all tarsomeres with erect bristles. Pulvilli shorter than last tarsal segment. Wing. Length: 2.4 mm. Upper side of basal costal cell with one long dark brown bristle. Fourth costal section 3 times as long as third costal section. Cross-vein r-m at middle of discal cell. 1–2 short dark setulae on tegula. Abdomen. Viewed obliquely from front tergites brown, tergite 1 silvery grey. Lateral bristles on first tergite present, 4–5 long dark bristles up to as long as hind femur’s width at base. Genitalia. Genital capsule in dorsal view: epandrium longer than wide (MLE: MWE = 1.3). widened at the base, elongated, moderately narrowed to apex, pointed, bent to each other at the apex ( Fig. 55A View FIGURE 55 ). Genital capsule in ventral view: both gonopods extended towards surstyli, pointed at apex with a distinct projection on inner margin; phallic guide with 3–4 short dorso- dorsolateral spines, one is longer; subepandrial sclerite widened, with a fin-shaped projection in its center, hypandrial apodeme distinct ( Fig. 55B View FIGURE 55 ); Genital capsule in lateral view: both surstyli straight in basal two thirds, then bent towards sternite in apical third ( Fig. 55D–E View FIGURE 55 ). Ejaculatory apodeme tube-like, bent, with a bulb in its middle ( Fig. 55C View FIGURE 55 ).

FEMALE: Unknown.

Etymology: This species is named after the type locality. Eight species of Tomosvaryella have been collected in this reserve and the only known specimen of this species is from there.

Examined material: HOLOTYPE: AUSTRALIA: New South Wales: Ledknapper National Reserve, 7.3km ENE Beulah Headquarters (LDN2M), 29°21’S 146°13’E, E. melanopholia , Spinifex , 10.XII.2009 – 18.III.2010, S. O’Sullivan, K. Taylor, Malaise trap, CNC576517 (1♂, QM).

Distribution: Australia (New South Wales) ( Fig. 128 View FIGURE 128 ).

Notes: Based on the shape of surstyli in dorsal view, this species is close to T. prolata sp. nov. It differs by the shape of surstyli, which are bent towards the sternites in the apical third in lateral view, with a fin-shaped projection in the middle of the subepandrial sclerite and a distinct hypandrial apodeme in ventral view ( Fig. 55A–E View FIGURE 55 ).